Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Subphylum Chelicerata Heymons, 1901
Euchelicerata Weygoldt & Paulus, 1979
Prosomapoda Lamsdell, 2013
Planaterga Lamsdell, 2013
Dekatriata Lamsdell, 2013
Sclerophorata Kamenz, Staude & Dunlop, 2011
Class Arachnida Lamarck, 1801
Micrura Hansen & Sørensen, 1904
Acaromorpha Dubinin, 1957
Subclass Acari Leach, 1817 - mites and ticks
Superorder Acariformes Zakhvatkin, 1952 - mitelike mites
Order Trombidiformes Reuter, 1909
Suborder Prostigmata Kramer, 1877
Infraorder Anystina van der Hammen, 1972
Hyporder Parasitengona Oudemans, 1909
Superfamily Trombiculoidea Ewing, 1929
Family Trombiculidaeᵀ Ewing, 1929 - chiggers
Subfamily Trombiculinaeᵀ Ewing, 1929
Tribe Trombiculiniᵀ Ewing, 1929
01.1 Genus Vatacarus Southcott, 1957
01.2 Genus Iguanacarus Vercammen-Grandjean, 1965
02 Genus Rhinibius Brennan & Yunker, 1969
03 Genus Brennanacarus Goff, Yunker & Wheeler, 1987
04 Genus Zumptrombicula Vercammen-Grandjean, 1967
05 Genus Myxacarus Brennan & Yunker, 1966
06 Genus Intercutestrix Brennan & Yunker, 1966
07 Genus Crotonasis Brennan & Yunker, 1966
08 Genus Blix Brennan & Yunker, 1966
09 Genus Fonsecula Loomis, 1966
10 Genus Tenotrombicula Vercammen-Grandjean, 1965
11 Genus Audytrombicula Vercammen-Grandjean, 1963
12 Genus Fereus Brennan & Jones, 1961
13 Genus Perissopalla Brennan & White, 1960
14 Genus Perates Brennan & Dalmat, 1960
15 Genus Multigniella Vercammen-Grandjean & Fain, 1957
16 Genus Speleocola Lipovsky, 1952
17 Genus Ipotrombicula Womersley, 1952
18 Genus Anomalaspis Brennan, 1952
19 Genus Pseudoblankaartia Fuller & Wharton, 1951
20.1 Genus Hexidionis Vercammen-Grandjean & Loomis, 1967
20.2 Genus Pentidionis Vercammen-Grandjean & Loomis, 1967, incertae sedis
21 Genus Eusaperium Brennan, 1970
22 Genus Atelepalme Brennan & Reed, 1973
23 Genus Carebareia Goff & Brennan, 1978
24 Genus Dolichotrombicula Feider, 1977
25 Genus Lacertacarus Shluger & Vasil'eva, 1977
26 Genus Nycterinastes Brennan & Reed, 1973
27 Genus Heterotectum Feider, 1983
28 Genus Otorhinophila Wrenn & Loomis, 1967
29 Genus Peltoculus Brennan, 1972
30 Genus Pentagonotectum Feider, 1983
31 Genus Tectumpilosum Feider, 1983
32 Genus Huabangsha Wen, Yu & Yang, 1980
33 Genus Hirsutiella Schluger & Vysotzkaya, 1970
34 Genus Cubanothrombium Feider, 1983
35 Genus Dongyangsha Wen, 1984
36 Genus Kepkatrombicula Kudryashova & Stekolnikov, 2010
37 Genus Muritrombicula Yu, Gong & Tao, 1981
38 Genus Octasternala Brown, 1990
39 Genus Paratrombicula Goff & Whitaker, 1984
40 Genus Phalcophila Brennan & Reed, 1973
41 Genus Poliremotus Brennan & Goff, 1978
42 Genus Striatiscuta Hsu & Hsu, 1982
43 Genus Teratothrix Brennan & Goff, 1978
44 Genus Xinjiangsha Wen & Shao, 1984
45.01 Genus Vanidicus Brennan & Jones, 1961
45.02 Genus Babiangia Southcott, 1954
45.03.1 Genus Trombiculaᵀ Berlese, 1905
45.03.2 Genus Boshkerria Fauran, 1959
45.03.3 Genus Grandjeana Koçak & Kemal, 2009
45.03.4 Genus Sasatrombicula Vercammen-Grandjean, 1960
45.03.5 Genus Rudnicula Vercammen-Grandjean, 1964
45.04 Genus Blankaartia Oudemans, 1911
45.05.1 Genus Oudemansidium Vercammen-Grandjean & André, 1966
45.05.2 Genus Willmannium Vercammen-Grandjean & Langston, 1976
45.05.3 Genus Chiroptella Vercammen-Grandjean, 1960
45.06.1.1 Genus Microtrombicula Ewing, 1950
45.06.1.2 Genus Marcandrea Vercammen-Grandjean, 1960
45.06.2.1 Genus Eutrombicula Ewing, 1938
45.06.2.2 Genus Blanciella Vercammen-Grandjean, 1960
45.07 Genus Polylopadium Brennan & Jones, 1961
45.08.1.1 Genus Tecomatlana Hoffmann, 1947
45.08.1.2 Genus Sauriscus Lawrence, 1949
45.08.2 Genus Beamerella Brennan, 1958
45.08.3 Genus Hooperella Vercammen-Grandjean, 1967
45.09 Genus Heaslipia Ewing, 1944
45.10 Genus Kaaia Brennan, 1958
45.11.1 Genus Leptotrombidium Nagayo, Miyagawa, Mitamura & Imamura, 1916
45.11.2 Genus Lorillatum Nadchatram, 1963
45.11.3.1 Genus Ericotrombidium Vercammen-Grandjean, 1965
45.11.3.2 Genus Toritrombicula Sasa, Hayashi & Kawashima, 1953
45.11.4 Genus Hypotrombidium Vercammen-Grandjean, 1960
45.12 Genus Miyatrombicula Sasa, Kawashima & Egashira, 1952
45.13 Genus Myotrombicula Womersley & Heaslip, 1943
45.14 Genus Whartonacarus Vercammen-Grandjean, 1960
45.15.1 Genus Fonsecia Radford, 1942
45.15.2 Genus Parasecia Loomis, 1966
45.16 Genus Neotrombicula Hirst, 1925
45.17 Genus Neotrombiculoides Vercammen-Grandjean, 1960
45.18 Genus Crotiscus Ewing, 1944
45.19 Genus Euschoengastoides Loomis, 1954
45.20 Genus Vercammenia Audy & Nadchatram, 1957
45.21 Genus Hoffmanniella Vercammen-Grandjean, 1960
45.22 Genus Aplodontophila Wrenn & Maser, 1981
45.23 Genus Oaxacarus Goff & Spicer, 1980
45.24 Genus Hyponeocula Vercammen-Grandjean, 1960
45.25 Genus Trombiculindus Radford, 1948
45.26 Genus Speotrombicula Ewing, 1946
46 Genus Afrotrombicula Kolebinova & Vercammen-Grandjean, 1978
47 Genus Ancoracarus Takahashi, Misumi & Takahashi, 2012
48 Genus Batmanacarus Bassini-Silva, Jacinavicius & Ochoa, 2021
49 Genus Bramkeria Bassini-Silva, Jacinavicius & Ochoa, 2021
50 Genus Brygoovia Stekolnikov & Fain, 2004
51 Genus Buclypeus Brennan, 1972
52 Genus Caamembecaia Gazêta, Amorim, Bossi, Linhares & Serra-Freire, 2006
53 Genus Hoffmannina Brennan & Jones, 1959
54 Genus Laotrombicula Stekolnikov, 2014
55 Genus Nahuacarus Bassini-Silva, Jacinavicius & Welbourn, 2020
56 Genus Novotrombicula Womersley & Kohls, 1947
57 Genus Pentagonaspis Vercammen-Grandjean & André, 1966
58 Genus Sambonacarus Stekolnikov, 2021
59 Genus Tanautarsala Brown, 2007
60 Genus Tauffliebiella Vercammen-Grandjean, 1960