Phylum Arthropoda Siebold, 1848 - arthropods
   Class Insecta Linnaeus, 1758 - insects
     Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Order Lepidoptera Linnaeus, 1758 - butterflies, moths
           Suborder Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Superfamily Papilionoidea Latreille, 1802
               Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                 Subfamily Nymphalinae Rafinesque, 1815
                   Section V see Tribe Coloburini
                     1 Genus Historis Hübner, 1819
                     2 Genus Smyrna Hübner, 1823
                     3 Genus Pycina Doubleday, 1849
                     4 Genus Baeotus Hemming, 1939
                     4.0 Genus Megistanis auct. see Genus Baeotus Hemming, 1939
                     5 Genus Colobura Billberg, 1820
                     5.0 Genus Gynaecia Doubleday, 1850 see Genus Colobura Billberg, 1820
                     6 Genus Tigridia Hübner, 1819
                     6.0 Genus Callizona Doubleday, [1848] see Genus Tigridia Hübner, 1819