Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade dorsates Cavalier-Smith, 2021
                   Clade podiates Cavalier-Smith, 2012
                     [crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                       [crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                         [crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                       Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                             Tactopoda Budd, 2001
                                               Phylum Arthropoda Siebold, 1848 - arthropods
                                                 Euarthropoda
                                                   Antennulata
                                                     Subphylum Mandibulata Snodgrass, 1938
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                         Altocrustacea
                                                           Miracrustacea
                                                             Superclass Hexapoda Latreille, 1825
                                                               Cercophora
                                                                 Class Insecta Linnaeus, 1758 - insects
                                                                   Subclass Dicondylia Haeckel, 1866
                                                                     Infraclass Pterygota Lang, 1888
                                                                       Metapterygota
                                                                         Neoptera Martynov, 1923
                                                                           Eumetabola
                                                                             Holometabola
                                                                               Order Coleoptera Linnaeus, 1758 - beetles
                                                                                 Suborder Polyphaga Emery, 1886
                                                                                   Series Scarabaeiformia Crowson, 1960
                                                                                     Superfamily Scarabaeoideaᵀ Latreille, 1802
                                                                                       Family Passalidae Leach, 1815
                                                                                         Subfamily Passalinaeᵀ Leach, 1815a
                                                                                           Tribe Macrolinini Kaup, 1871
                                                                                             01 Genus Macrolinusᵀ Kaup, 1868a
                                                                                             02 Genus Aceraius Kaup, 1868
                                                                                             03 Genus Gonatas Kaup, 1871
                                                                                             04 Genus Mastachilus Kaup, 1868a
                                                                                             05 Genus Pharochilus Kaup, 1868a
                                                                                             06 Genus Tarquinius Kuwert, 1891
                                                                                             07 Genus Kaupiolus Zang, 1903
                                                                                             08 Genus Aurelius Kuwert, 1891
                                                                                             09 Genus Chilomazus Zang, 1905
                                                                                             10 Genus Pelopides Kuwert, 1896
                                                                                             11 Genus Pleurarius Kaup, 1868b
                                                                                             12 Genus Austropassalus Mjöberg, 1917