Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Hydrophiloidea Latreille, 1802
Family Hydrophilidaeᵀ Latreille, 1802 - water scavenger beetles
Genus Agna Smetana, 1978
Genus Ametor Semenov, 1900
Genus Dactylosternum Wollaston, 1854
Genus Derallus Sharp, 1882
Genus Dibolocelus Regimbart, 1901
Genus Epimetopus Lacordaire, 1854
Genus Genyon Smetana, 1978
Genus Georissus Latreille, 1809
Genus Helobata Bergroth, 1888
Genus Helocombus Horn, 1890
Genus Helophorus Fabricius, 1775
Genus Hydrobiomorpha Blackburn, 1888
Genus Hydrochus Leach, 1817
Genus Ictenosternum Smetana, 1978
Genus Omicrus Sharp, 1879
Genus Oosternum Sharp, 1882
Genus Pelosoma Mulsant, 1844
Genus Pemelus Horn, 1890
Genus Phaenonotum Sharp, 1882
Genus Spercheus Kugelann, in Illiger, 1798
Genus Sperchopsis LeConte, 1862
Genus Tropisternus Solier, 1834
1 Subfamily Sphaeridiinae (Latreille, 1802)
2 Subfamily Hydrophilinaeᵀ Latreille, 1802