Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Staphylinoideaᵀ Latreille, 1802
Family Leiodidae Fleming, 1821
Genus Adelopsis Portevin, 1907
Genus Aglyptinus Cockerell, 1906
Genus Anogdus LeConte, 1866
Genus Apheloplastus Brown, 1937
Genus Caenocyrta Brown, 1937
Genus Cainosternum Notman, 1921
Genus Caloptrichus Murry, 1856
Genus Catopocerus Motschulsky, 1869
Genus Colon Herbst, 1797
Genus Dissochaetus Reitter, 1885
Genus Ecarinosphaerula Hatch, 1929
Genus Echinocoleus Horn, 1885
Genus Glacicavicola Westcott, 1968
Genus Isoplastus Horn, 1880
Genus Lionothus Brown, 1937
Genus Neocyrtusa Brown, 1937
Genus Platycholeus Horn, 1880
Genus Prionochaeta Horn, 1880
Genus Stetholiodes Fall, 1910
1 Subfamily Ptomaphaginae
2 Subfamily Nemadinae
3 Subfamily Anemadinae
4.1 Subfamily Cholevinae (Kirby, 1837)
4.2 Subfamily Catopinae
5 Subfamily Bathysciinae
6 Subfamily Leiodinaeᵀ Fleming, 1821
7.1 Subfamily Leptininae LeConte, 1866
7.2 Subfamily Platypsyllinae Ritsema, 1869