Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Staphylinoideaᵀ Latreille, 1802
Family Staphylinidaeᵀ Latreille, 1802 - rove beetles
Genus Abdiunguis Park & Wagner, 1962
Genus Acalophaena Sharp, 1886
Genus Achenomorphus Motschoulsky, 1858
Genus Acolonia Casey, 1893
Genus Acrimea Casey, 1911
Genus Acrostilicus Hubbard, 1896
Genus Acrotona Thomson, 1859
Genus Actiastes Casey, 1897
Genus Actium Casey, 1887
Genus Actizona Chandler, 1985
Genus Adinopsis Cameron, 1918
Genus Adota Casey, 1910
Genus Adranes Le Conte, 1850
Genus Agaricomorpha Ashe, 1984
Genus Aidochara Casey, 1906
Genus Alapsodus Tottenham, 1939
Subfamily Aleocharomorphes
Genus Aleodorus Say, 1839
Genus Alisalia Casey, 1911
Genus Allobrox Fletcher, 1928
Genus Allotrimium Park, 1943
Genus Amblopusa Casey, 1893
Genus Amphicroum Kraatz, 1858
Genus Anacyptus Horn, 1877
Genus Anaduosternum Notman, 1922
Genus Anaquedius Casey, 1915
Genus Anatheta Casey, 1910
Genus Anchylarthron Brendel, 1887
Genus Anepsiota Casey, 1893
Genus Anitra Casey, 1893
Genus Anotylus Thomson, 1859
Genus Apalonia Casey, 1906
Genus Apheloglossa Casey, 1893
Genus Apocellus Erichson, 1839
Genus Apteronina Wasmann, 1901
Genus Araeocerus Nordmann, 1837
Genus Arianops Brendel, 1893
Genus Arthmius Le Conte, 1850
Genus Artochia Casey, 1893
Genus Asemobius Horn, 1895
Genus Asthenesita Casey, 1893
Genus Athetalia Casey, 1910
Genus Athetota Casey, 1906
Genus Atinus Horn, 1868
Genus Baeocera Erichson, 1845
Genus Bamona Sharp, 1883
Genus Basolum Casey, 1897
Genus Batriasymmodes Park, 1951
Genus Beeria Hatch, 1957
Genus Belonuchus Nordmann, 1837
Genus Bessobia Lohse, 1990
Genus Beyeria Fenyes, 1910
Genus Biocrypta Casey, 1905
Genus Biotus Casey, 1887
Genus "Bisnius" Leach, 1829, nom. nud.
Genus Blepharhymenus Solier, 1849
Genus Bolitopunctus Campbell, 1993
Genus Bontomtes Grigarick & Schuster, 1980
Genus Boreostiba Lohse, 1990
Genus Brathinus LeConte, 1852
Genus Briaraxis Brendel, 1894
Genus Bryobiota Casey, 1893
Genus Bryophacis Reitter, 1909
Genus Bryothinusa Casey, 1904
Genus Bythinoplectus Reitter, 1882
Genus Caccoplectus Sharp, 1887
Genus Cainotyphlus Coiffait, 1962
Genus Canastota Casey, 1910
Genus Carphacis Des Gozis, 1886
Genus Cedius Leconte, 1850
Genus Ceophyllus LeConte, 1850
Genus Cercocerus LeConte, 1861
Genus Charhyphus Sharp, 1887
Genus Chionotyphlus Smetana, 1986
Genus Clavilispinus Bernhauer, 1926
Genus Clusiota Casey, 1910
Genus Conoplectus Brendel, 1888
Genus Crephalia Casey, 1910
Genus Crinolinus Smetana, 1982
Genus Ctenisis Raffray, 1890
Genus Cupila Casey, 1897
Genus Custotychus Park & Wagner, 1961
Genus Cylindrarctus Schaufuss, 1887
Genus Cyparium Erichson, 1845
Genus Cyphea Fauvel, 1863
Genus Dacnochilus LeConte, 1861
Genus Dalmodes Reitter, 1882
Genus Dalmosella Casey, 1897
Genus Datomicra Mulsant & Rey, 1873
Genus Decarthron Brendel, 1865
Genus Decusa Casey, 1900
Genus Devia Blackwelder, 1952
Genus Dexiogyia Thomson, 1858
Genus Diaulota Casey, 1893
Genus Dimetrota Mulsant & Rey, 1873
Genus Dinardilla Wasmann, 1901
Genus Dinocoryna Casey, 1893
Genus Diochus Erichson, 1839
Genus Doliponta Blackwelder, 1952
Genus Dytoscotes Smetana, 1980
Genus Earota Mulsant & Rey, 1874
Genus Eburniogaster Seevers, 1938
Genus Ecbletus Sharp, 1887
Genus Echiaster Erichson, 1840
Genus Ecitocala Frank & Thomas, 1981
Genus Ecitonidia Wasmann, 1900
Genus Ecitoxenida Wasmann, 1909
Genus Elachistarthron Notman, 1920
Genus Eleusis Laporte, 1835
Genus Empelus LeConte, 1862
Genus Ephelinus Cockerell, 1906
Genus Euboarhexius Grigarick & Schuster, 1966
Genus Eudectoides Campbell, 1978
Genus Eumicrota Casey, 1906
Genus Euphalepus Reitter, 1883
Genus Euplecturga Park & Wagner, 1961
Genus Eupsenius LeConte, 1850
Genus Euromota Casey, 1906
Genus Euthorax Solier, 1849
Genus Eutrichites LeConte, 1880
Genus Eutyphlus LeConte, 1880
Genus Euvira Sharp, 1883
Genus Falagrioma Casey, 1906
Genus Falagriota Casey, 1906
Genus Fenderia Hatch, 1957
Genus Foveoscapha Park & Wagner, 1961
Genus Fusalia Casey, 1911
Genus Fustiger Brendel, 1866
Genus Gaenima Casey, 1911
Genus Gnathoryphium Campbell, 1978
Genus Gnathusa Fenyes, 1909
Genus Gnypetella Casey, 1906
Genus Goniusa Casey, 1906
Genus Gyronycha Casey, 1893
Genus Habrolinus Casey, 1906
Genus Hadrotes Mäklin, 1852
Genus Haida Keen, 1897
Genus Hamotus Aube, 1844
Genus Hatchia Park & Wagner, 1961
Genus Hemiquedius Casey, 1915
Genus Hesperolinus Casey, 1906
Genus Hesperotychus Schuster & Marsh, 1958
Genus Heteroleptotyphlus Coiffait, 1962
Genus Holoboreaphilus Campbell, 1978
Genus Homaeotarsus Hochhuth, 1851
Genus Homalotusa Casey, 1906
Genus Homeotyphlus Coiffait, 1962
Genus Hoplandria Kraatz, 1857
Genus Hypnogyra Casey, 1906
Genus Hypotelus Erichson, 1839
Genus Iotota Casey, 1910
Genus "Ischnosoma" Stephens, 1829, nom. nud.
Genus Laetulonthus Moore & Legner, 1972
Genus Lamiota Casey, 1910
Genus Leistotrophus Perty, 1830
Genus Lemelba Park, 1953
Genus Lepitacnus Smetana, 1982
Genus Leptagria Casey, 1906
Genus Leptobamona Casey, 1911
Genus Leptochirus Germar, 1824
Genus Leptoplectus Casey, 1908
Genus Linohesperus Smetana, 1982
Genus Liparocephalus Maeklin, 1853
Genus Lispinus Erichson, 1839
Genus Lissagria Casey, 1906
Genus Lissobiops Casey, 1905
Genus Lissohypnus Casey, 1906
Genus Lithocharodes Sharp, 1876
Genus Lobrathium Mulsant & Rey, 1878
Genus Longipeltina Bernhauer, 1912
Genus Lordithon Thomson, 1859
Genus Losiusa Seevers, 1978
Genus Lucifotychus Park & Wagner, 1961
Genus Lypoglossa Fenyes, 1918
Genus Machaerites Muller, 1855
Genus Machaerodes Brendel, 1890
Genus Mathrilaeum Moore, 1966
Genus Mayetia Mulsant & Rey, 1875
Genus Megalopinus Eichelbaum, 1915
Genus Megarafonus Casey, 1897
Genus Megastilicus Casey, 1889
Genus Melanalia Casey, 1911
Genus Melba Casey, 1897
Genus Meronera Sharp, 1887
Genus Micratheta Casey, 1910
Genus Micrearota Casey, 1910
Genus Microbledius Herman, 1972
Genus Microdonia Casey, 1893
Genus Microdota Mulsant & Rey, 1873
Genus Microedus LeConte, 1874
Genus Microlinus Casey, 1906
Genus Mipselytrus Park, 1953
Genus Mitosynum Campbell, 1982
Genus Mocyta Mulsant & Rey, 1874
Genus Moluciba Casey, 1911
Genus Morius Casey, 1893
Genus Myrmecocephalus MacLeay, 1873
Genus Myrmobiota Casey, 1893
Genus Nacaeus Blackwelder, 1942
Genus Nanobius Herman, 1977
Genus Nearctitychus Chandler, 1988
Genus Neobeyeria Jacobson et al., 1987
Genus Neobolitobius Campbell, 1993
Genus Neohypnus Coiffait & Saiz, 1964
Genus Neoleptotyphlus Coiffait, 1959
Genus Neomedon Sharp, 1886
Genus Neotrochus Blackwelder, 1943
Genus Neoxantholinus Cameron, 1904
Genus Neoxus Herman, 1970
Genus Nisaxis Casey, 1886
Genus Nitidotachinus Campbell, 1993
Genus Nordenskioeldia Sahlberg, 1880
Genus Nosora Casey, 1911
Genus Noverota Casey, 1910
Genus Ochthephilum Stephens, 1829
Genus Ochthephilus Mulsant & Rey, 1856
Genus Omalonomus Campbell & Peck, 1990
Genus Omalorphanus Campbell & Chandler, 1987
Genus Omegalia Casey, 1910
Genus Ophioomma Notman, 1920
Genus Orobanus LeConte, 1878
Genus Oropodes Casey, 1893
Genus Oropus Casey, 1886
Genus Orthodiatelus Notman, 1920
Genus Orus Casey, 1884
Genus Osorius Latreille, 1829
Genus Ouachitychus Chandler, 1988
Genus Oxybleptes Smetana, 1982
Genus Pachycerota Casey, 1906
Genus Pachystilicus Casey, 1905
Genus Paederopsis Wasman, 1912
Genus Palaminus Erichson, 1839
Genus Panalota Casey, 1910
Genus Pancota Casey, 1906
Genus Paradilacra Bernhauer, 1890
Genus Parameotica Ganglbauer, 1895
Genus Parocalea Bernhauer, 1901
Genus Parothius Casey, 1906
Genus Pelecomalium Casey, 1886
Genus Pentanota Bernhauer, 1905
Genus Phanerota Casey, 1906
Genus Phasmota Casey, 1910
Genus Philhygra Mulsant & Rey, 1873
Genus Philotermes Kraatz, 1857
Genus Phlaeopterus Motschulsky, 1852
Genus Piestus Gravenhorst, 1806
Genus Pilactium Grigarick & Schuster, 1970
Genus Pilopius Casey, 1897
Genus Pinophilus Gravenhorst, 1802
Genus Platandria Casey, 1893
Genus Platyprosopus Mannerheim, 1831
Genus Pontomalota Casey, 1885
Genus Prespelea Park, 1953
Genus Probeyeria Seevers, 1965
Genus Prototyphlus Coiffait, 1962
Genus Psamathobledius Herman, 1972
Genus Pselaptrichus Brendel, 1899
Genus Pselaptus LeConte, 1880
Genus Psephidonus Gistel, 1856
Genus Pseudactium Casey, 1908
Genus Pseudocypus Mulsant & Rey, 1876
Genus Pseudomegista Bernhauer, 1907
Genus Pseudota Casey, 1910
Genus Pseudotrimium Raffray, 1898
Genus Pseudousipalia Lohse, 1990
Genus Pulicomorpha Mann, 1924
Genus Pygmactium Schuster & Grigarick, 1968
Genus Rabigus Mulsant & Rey, 1876
Genus Ramecia Casey, 1893
Genus Renardia Motschulsky, 1865
Genus Rhexidius Casey, 1887
Genus Rhexius LeConte, 1850
Genus Rhinoscepsis LeConte, 1878
Genus Rhybaxis Saulcy, 1874
Genus Rimulincola Sanderson, 1947
Genus Rugilus Leach, 1819
Genus Sableta Casey, 1910
Genus Salinamexus Moore & Legner, 1977
Genus Sauridus Mulsant & Rey, 1876
Genus Saurohypnus Sharp, 1885
Genus Saxet Grigarick & Schuster, 1980
Genus Scalenarthrus LeConte, 1880
Genus Schistacme Notman, 1920
Genus Seeversiella Ashe, 1986
Genus Sepedophilus Gistel, 1856
Genus Sibiota Casey, 1906
Genus Silusida Casey, 1906
Genus Simplona Casey, 1897
Genus Sipaliella Casey, 1911
Genus Sonoma Casey, 1886
Genus Speleobama Park, 1951
Genus Stamnoderus Sharp, 1886
Subfamily Staphylinomorphes
Genus Stenistoderus Jaquelin du Val, 1856
Genus Stethusa Casey, 1910
Genus Stictocranius LeConte, 1866
Genus Stictolinus Casey, 1906
Genus Stilicolina Casey, 1905
Genus Stilicopsis Sachse, 1852
Genus Stilomedon Sharp, 1886
Genus Stratus Schaufuss, 1872
Genus Strigota Casey, 1910
Genus Strophogastra Fenyes, 1921
Genus Subhaida Hatch, 1957
Genus Sunius Curtis, 1829
Genus Symbiochara Fenyes, 1909
Genus Synaptina Casey, 1910
Genus Tachinomorphus Kraatz, 1859
Genus Tanyrhinus Mannerheim, 1852
Genus Tarphiota Casey, 1893
Genus Teliusa Casey, 1906
Genus Telotyphlus Coiffait, 1962
Genus Termitomimus Trägårdh, 1907
Genus Termitonidia Seevers, 1938
Genus Teropalpus Solier, 1849
Genus Tetradonia Wasmann, 1894
Genus Tetralaucopora Bernhauer, 1928
Genus Tetrallus Bernhauer, 1905
Genus Tetrascapha Schuster & Marsh, 1957
Genus Texamaurops Barr & Steeves, 1963
Genus Thecturota Casey, 1893
Genus Thesiastes Casey, 1893
Genus Thesium Casey, 1884
Genus Thinocharis Kraatz, 1859
Genus Thinodromus Kraatz, 1858
Genus Thinopinus LeConte, 1852
Genus Thinusa Casey, 1893
Genus Thyasophila Fairmaire & Laboulbene, 1856
Genus Thyreocephalus Guerin-Meneville, 1844
Genus Timagenes Smetana, 1982
Genus Tmesiphorus LeConte, 1849
Genus Tomoplectus Raffray, 1898
Genus Toxidium LeConte, 1860
Genus Trachyota Casey, 1906
Genus Trichiusa Casey, 1893
Genus Trichopsenius Horn, 1877
Genus Trigonodemus LeConte, 1863
Genus Trigonoplectus Bowman, 1934
Genus Trigonopselaphus Gemminger & Harold, 1868
Genus Trigonurus Mulsant, 1847
Genus Trimioarcus Park, 1952
Genus Trimiomelba Casey, 1897
Genus Trimioplectus Brendel, 1890
Genus Trisignis Park & Schuster, 1955
Genus Tympanophorus Nordmann, 1837
Genus Unamis Casey, 1893
Genus Valda Casey, 1893
Genus Valenusa Casey, 1906
Genus Vellejus Mannerheim, 1830
Genus Vellica Casey, 1885
Genus Vicelva Moore & Legner, 1973
Genus Xanthopygus Kraatz, 1857
Genus Xenicopoda Moore & Legner, 1971
Genus Xenistusa LeConte, 1880
Genus Xenomedon Fall, 1912
Genus Xenota Mulsant & Rey, 1874
Genus Xenotyphlus Coiffait, 1962
Genus Xestolinus Casey, 1906
Genus Xesturida Casey, 1906
Genus Zalobius LeConte, 1874
Genus Zenon Smetana, 1982
Genus Zonaira Grigarick & Schuster, 1980
Subfamily Pselaphinaeᵀ (Latreille, 1802)
01 Subfamily Micropeplinae (Leach, 1815)
02 Subfamily Scaphidiinae Latreille, 1807
03 Subfamily Piestinae (Erichson, 1839)
04 Subfamily Osoriinae Erichson, 1839
05 Subfamily Pseudopsinae Ganglbauer, 1895
06 Subfamily Phloeocharinae (Erichson, 1839)
07 Subfamily Olisthaerinae Thomson, 1858
08 Subfamily Metopsiinae
09 Subfamily Proteininae Erichson, 1839
10 Subfamily Omaliinae MacLeay, 1825
11 Subfamily Oxytelinae (Fleming, 1821)
12 Subfamily Leptotyphlinae Fauvel, 1874
13 Subfamily Oxyporinae (Fleming, 1821)
14 Subfamily Steninae MacLeay, 1825
16 Subfamily Euaesthetinae Thomson, 1859
17 Subfamily Paederinae (Fleming, 1821)
18 Subfamily Xantholininae Erichson, 1839
19 Subfamily Staphylininaeᵀ Latreille, 1802
20 Subfamily Habrocerinae Mulsant & Rey, 1877
21 Subfamily Trichophyinae Thomson, 1858
22 Subfamily Tachyporinae MacLeay, 1825
23 Subfamily Hypocyphtinae
24 Subfamily Aleocharinae (Fleming, 1821)
25 Subfamily Termitodiscinae
26 Subfamily Pygosteninae
27 Subfamily Dasycerinae Reitter, 1887