Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade natates Cavalier-Smith, 2021
                   Clade photaria Cavalier-Smith, 2021
                     Clade eucorta Cavalier-Smith, 2021
                       Superkingdom Corticata Lankester, 1878 - corticates
                         Kingdom Chromista Caval.-Sm. (1981) - chromists
                           Subkingdom Harosa Cavalier-Smith, 2010
                             [crown] Clade Sar Adl et al. 2012 [Simpson & Dunthorn 2020]
                               Infrakingdom Halvaria Cavalier-Smith, 2010
                                 Superphylum Alveolata Cavalier-Smith, 1991
                                   Phylum Ciliophora Doflein, 1901 - ciliates
                                     Subphylum Intramacronucleata Lynn, 1996 - amphiesmates
                                       Class Colpodea Small & Lynn, 1981
                                         Order Colpodidaᵀ de Puytorac et al., 1974
                                           Family Grandoriidae Corliss, 1960
                                             Genus Grandoriaᵀ Corliss, 1960
                                               Grandoria aculeataᵀ (Grandori & Grandori, 1934) Corliss, 1960