Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade natates Cavalier-Smith, 2021
                   Clade photaria Cavalier-Smith, 2021
                     Clade eucorta Cavalier-Smith, 2021
                       Superkingdom Corticata Lankester, 1878 - corticates
                         Kingdom Plantae Haeckel (1866) - plants
                           Subkingdom Viridiplantae Caval.-Sm. (1981) - green plants
                             Infrakingdom Streptophyta Caval.-Sm. (1993)
                               [crown] Clade Phragmoplastophyta Lecointre & Le Guyader 2006 [Hall, McCourt, Delwiche, Mishler & Karol 2020]
                                 [crown] Clade Embryophytaᵀ Engler 1887 [Mishler & Qiu 2020]
                                   Phylum Bryophytaᵀ A.Braun (1864) - bryophytes
                                     [crown] Clade "Setaphyta" Puttick, Morris, Williams, Cox, Edwards, Kenrick, Pressel, Wellman, Schneider, Pisani & Donoghue 2018
                                       Subphylum Marchantiophytina Doweld (2001) - liverworts
                                         Class Jungermanniopsida Stotler & Crand.-Stotl. (1977) - leafy liverworts
                                           Subclass Jungermanniidaeᵀ Engl. (1893)
                                             Order Jungermannialesᵀ H. Klinggr.
                                               Suborder Lophocoleineae Schljakov
                                                 Family Lophocoleaceaeᵀ Vanden Berghen
                                                   01 Genus Amphilophocolea R.M.Schust. (2001)
                                                   02 Genus Chiloscyphus Corda (1829), nom. cons., orth. cons.
                                                   03 Genus Clasmatocolea Spruce (1885)
                                                   04 Genus Conoscyphus Mitt.
                                                   05 Genus Cyanolophocolea R.M. Schust.
                                                   06 Genus Evansianthus R.M. Schust. & J.J. Engel (1973)
                                                   07 Genus Hepatostolonophora J.J. Engel & R.M. Schust.
                                                   08 Genus Heteroscyphus Schiffn.
                                                   09 Genus Lamellocolea J.J. Engel
                                                   10 Genus Leptophyllopsis R.M. Schust. (1963)
                                                   11 Genus Leptoscyphopsis R.M. Schust.
                                                   12 Genus Leptoscyphus Mitt. (1851)
                                                   13 Genus Lophocoleaᵀ (Dumort.) Dumort. (1835)
                                                   14 Genus Pachyglossa Herzog & Grolle (1959)
                                                   15 Genus Perdusenia Hässel
                                                   16 Genus Physotheca J.J. Engel & Gradst.
                                                   17 Genus Pigafettoa C. Massal.
                                                   18 Genus Platycaulis R.M. Schust.
                                                   19 Genus Pseudolophocolea R.M.Schust. & J.J. Engel (1982)
                                                   20 Genus Stolonivector J.J. Engel
                                                   21 Genus Xenocephalozia R.M. Schust. (1965)