Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 Clade neozoa Cavalier-Smith 1993
                   Clade scotokaryotes Cavalier-Smith 2000
                     Clade podiates Cavalier-Smith 2012
                       [crown] Clade Amorphea Adl et al. 2012
                         [crown] Clade Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
                             Kingdom Fungi R.T.Moore (1980) - fungi
                               Subkingdom Dikarya Hibbett, T.Y. James & Vilgalys (2007)
                                 Phylum Basidiomycota R.T.Moore (1980) - basidiomycetes
                                   Subphylum Ustilaginomycotina Doweld (2001)
                                     1 Class Exobasidiomycetes Begerow, M. Stoll & R. Bauer (2007) [paraphyletic]
                                     1.n1.1 Anamorph class Malasseziomycetes Denchev & T. Denchev (2014) [monotypic]
                                     1.n1.2 Anamorph class Moniliellomycetes Q.M. Wang, F.Y. Bai & Boekhout (2014) [monotypic]
                                     1.n2 Class Ustilaginomycetesᵀ Warm. (1884) - smut fungi