Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum †Artiopoda Hou & Bergström, 1997
Superclass †Trilobitomorpha Størmer, 1944
Class †Trilobita Walch, 1771 - trilobites
Genus †Abakanopleura Sivov, 1955
Genus †Abdulinaspis Ergaliev, 1980
Genus †Acanthocephalus Qiu, 1983, nom. illeg.
Genus †Acantholenus Matthew, 1898
Genus †Acanthomicmacca Hupé, 1953
Genus †Acrohybus Raymond, 1937
Genus †Adelogonus Opik, 1967
Genus †Aethia Qian & Zhou, 1984, nom. illeg.
Genus †"Aglagalia" Hupé, 1960, nom. nud.
Genus †Agraulaeva
Genus †Aidarella Lermontova, 1951
Genus †Aktugaiella Ergaliev, 1980
Genus †Alomataspis Opik, 1967
Genus †Altajaharpes Petrunina, 1984
Genus †Amechilus Ross, 1951
Genus †Americare Lochman & Hu, 1960
Genus †Amgaspis Chernysheva, 1956
Genus †Amginouyia Chernysheva, 1956
Genus †Ampellatocephalina Lu & Qian, 1983
Genus †Anacopodia
Genus †Antagmopleura Lermontova, 1960
Genus †Antatlaeva Hupé, 1960
Genus †Asaphelus
Genus †Ascetopeltis Owens, 1973
Genus †Atratebia Shergold, 1975
Genus †Barklyella Shergold, 1969
Genus †Beikedamaspis
Genus †"Bergerolimbus" Hupé, 1960, nom. nud.
Genus †Bicornipyge Lisogor, 1961
Genus †Bienvillia Clark, 1924
Genus †Blayacina Cobbold, 1932
Genus †Bodenbenderia Harrington & Leanza, 1957
Genus †Bondarevites Lazarenko, 1989
Genus †Brackebuschia Harrington & Leanza, 1957
Genus †Brevitermierella Hupé, 1953
Genus †Bridgeia Lochman, 1944
Genus †Cambrunicornia Geyer, 1990
Genus †Canningella Legg, 1976
Genus †Carmon Barrande, 1872
Genus †Catadoxides Matthew, 1899
Genus †Cermataspis Opik, 1967
Genus †Cermatops Shergold, 1980
Genus †Chalfontia Shergold, 1982
Genus †Chekiangaspis Lu, 1961
Genus †Chinensis
Genus †Chittidilla King, 1941
Genus †Chondroparia Lorenz, 1906
Genus †Conoides Howell, 1937
Genus †Conopolus Robison, 1988
Genus †Cornovica Whittard, 1960
Genus †Coronocephalus Grabau, 1924
Genus †Costonia Whittard, 1956
Genus †Crossoura Moberg & Segerberg, 1906
Genus †Crucicephalus Shergold, 1971
Genus †Cryptoderaspis Rasetti, 1946
Genus †Curiaspis Sdzuy, 1955
Genus †Cuyanaspis Rusconi, 1953
Genus †Cyclolenzellina
Genus †Deltacare Harrington & Leanza, 1957
Genus †Dicanthopyge Palmer, 1965
Genus †Durinia Robison, 1988
Genus †Edganocare
Genus †Eilura Resser & Endo, 1935
Genus †Elburgia Palmer, 1960
Genus †"Ellipsuella" Hupé, 1960, nom. nud.
Genus †Ellsaspis Rasetti, 1945
Genus †Endoaspis Lochman, 1956
Genus †Entsyna Rozova, 1968
Genus †Eosaukia Lu, 1954
Genus †Esseigania Kobayashi, 1944
Genus †Eulomella Kobayashi, 1955
Genus †Fabulaspis Ivshin, 1952
Genus †Folliceps Opik, 1970
Genus †Galahetes Opik, 1975
Genus †Gentilaspis Hupé, 1958
Genus †Gignopeltis Raymond, 1924
Genus †Gigoutella Hupé, 1953
Genus †Gogoella Legg, 1976
Genus †Goycoia Rusconi, 1950
Genus †Hamashania Kobayashi, 1941
Genus †Hercantyx Öpik, 1963
Genus †Hicksia Delgado, 1904
Genus †Holyoakia Palmer, 1995
Genus †Hongshiyanaspis Zhang & Lin, 1980
Genus †"Howelluella" Hupé, 1960, nom. nud.
Genus †Huamiaocephalus Qiu, 1983
Genus †Ishpella Petrunina, 1984
Genus †Issafenia Hupé, 1960
Genus †Issafeniella Geyer, 1990
Genus †Iveria Shergold, 1980
Genus †Iwayaspis Kobayashi, 1962
Genus †Jegorovaia Lu, 1964
Genus †Jonotus Meyer, 1848
Genus †Jubileia Kobayashi, 1938
Genus †Juriietella Jegorova, 1981
Genus †Kamlacus Petrunina, 1984
Genus †Kazellina Sivov, 1955
Genus †Kerbinella Jegorova, 1984
Genus †Knechtelia Lochman, 1950
Genus †Koldinia Walcott & Resser, 1928
Genus †Koldiniella Sivova, 1955
Genus †Kolpura Resser & Endo, 1935
Genus †Koraipsis Kobayashi, 1934
Genus †Kujandina
Genus †Kunmingaspis Lu & Chang, 1974
Genus †Kunshanaspis Zhang & Zhou, 1985
Genus †Kymataspis Geyer, 1990
Genus †Latikingaspis Geyer, 1990
Genus †Latoucheia Hupé, 1953
Genus †Leichneyella Opik, 1967
Genus †Leptopilus Raymond, 1924
Genus †Levisia Walcott, 1911
Genus †Liaoyangaspis Chang, 1959
Genus †Liriamnica Shergold, 1980
Genus †Lomsucaspis
Genus †Lotosoides Shergold, 1975
Genus †Loxonepea Opik, 1970
Genus †Luojiashania Chang, 1974
Genus †Lusampa Petrunina, 1984
Genus †Macrogrammus Whittard, 1966
Genus †Mannschreckia
Genus †Marcaisia
Genus †Mareda Kobayashi, 1942, nom. illeg.
Genus †Matania Rasetti, 1946
Genus †Mataninella Sivor, 1955
Genus †Mecophrys Shergold, 1982
Genus †Megagraulos Kobayashi, 1935
Genus †Megalaspidicus Petrunina, 1984
Genus †Megalopsis Qian, 1983
Genus †Meisterella Ivshin, 1952
Genus †Mendosina Shergold, 1975
Genus †Meropalla Opik, 1967
Genus †Metagraulos Kobayashi, 1935
Genus †Moretarva
Genus †Nannopeltis Harrington & Leanza, 1957
Genus †Ninaspis Ivshin, 1953
Genus †Norina
Genus †Notopeltis Harrington & Leanza, 1957
Genus †Oligometopus Resser, 1936
Genus †Onchonotus Raymond, 1924
Genus †Opipeuter Fortey, 1974, nom. illeg.
Genus †Orictocara
Genus †Orienturus Repina, 1973
Genus †Ornamentaspis Geyer, 1990
Genus †Orodes Geyer, 1990, nom. illeg.
Genus †Otenaspella
Genus †Ourikaia Geyer, 1990
Genus †Oxygonaspis Zhang, 1983
Genus †Paichiashania
Genus †Palaeolentus
Genus †Palmeraspis Repina, 1973
Genus †Parabolinella Brögger, 1882
Genus †Paraplagiura Zhang, 1983
Genus †Paraplesiograulos
Genus †Pararaia Kobayashi, 1942
Genus †Pararania
Genus †Peculicephalina Qian Yiyuan, 1985
Genus †Penarosa Opik, 1970
Genus †Pensacola Palmer & Gatehouse, 1972, nom. illeg.
Genus †Phoreotropis Raymond, 1924
Genus †Phylacterus Raymond, 1924
Genus †Piliolites Cozzens, 1846
Genus †Plagiura Resser, 1935
Genus †Planolimbus Geyer, 1990
Genus †Poculicephalina Qian, 1986
Genus †Priceaspis Legg, 1976
Genus †Probilacunaspis Ergaliev, 1980
Genus †Prohedinella Sivov, 1955
Genus †Pseudoacicaudus
Genus †Pseudoclelanida
Genus †Pseudocrecephalaspina
Genus †Pseudoliostracina Kobayashi, 1938
Genus †Pseudosalteria Raymond, 1924
Genus †Pseudosaukia Rasetti, 1944
Genus †"Pseudotermierella" Hupé, 1960, nom. nud.
Genus †Ptyttstrigis Hohensee, 1989
Genus †Pyraustocranium Ross, 1951
Genus †Qijiangia Xiang & Ji, 1987
Genus †Redlichaspis Kobayashi, 1935
Genus †Resseria Sun, 1937
Genus †Resseropsis Hupé, 1960
Genus †"Rochaeva" Hupé, 1960, nom. nud.
Genus †Rongxiella Chang, 1974
Genus †Rowia Deiss, 1939
Genus †Sarkia Koucek, 1919
Genus †Schistocephalus Chernysheva, 1956, nom. illeg.
Genus †Schistometopus Resser, 1938
Genus †Sectigena Geyer, 1990
Genus †Senticucullus Xia, MS
Genus †Serksia Hupé, 1960
Genus †Serksioides Hupé, 1960
Genus †Shipaiella Zhang & Qian, 1980
Genus †Sibiriopeltis Petrunina, 1984
Genus †Sinolenus Kobayashi, 1944
Genus †Smeathenia Dean, 1960
Genus †Sphaerocare Harrington & Leanza, 1957
Genus †Spirantyx Whittard, 1961
Genus †Strigambitus Palmer, 1965
Genus †Sunaspis Lu, 1953
Genus †Tadakoustia Geyer, 1990
Genus †Taidonorus
Genus †Tamdaspis Lisogor, 1977
Genus †Taplimella
Genus †Tasmanaspis Kobayashi, 1940
Genus †Tazzemmourtia
Genus †Tholifrons Palmer, 1968
Genus †Thomondia Harper, 1942
Genus †Thoracocare Robison & Campbell, 1974
Genus †Thymurus Whittard, 1966
Genus †Tongxingaspis
Genus †Townleyella Opik, 1967
Genus †Toxotis Wallerius, 1895
Genus †Trinia Poletaeva, 1956
Genus †Ullaspis Westergard, 1948
Genus †Ulrickaspis
Genus †Validaspis Repina, 1977
Genus †Verditerrina Robison, 1988
Genus †Volonellus Ivshin, 1952
Genus †Wandelella Robison, 1988
Genus †Westonaspis Rasetti, 1945
Genus †Whittardolithus Hughes, Ingham & Addison, 1975
Genus †Winsteria
Genus †Xystridura Whitehouse, 1936
Genus †Yunnanocephalus Kobayshi, 1936
Genus †Zacompsus Raymond, 1924
Genus †"Zigzuella" Hupé, 1960, nom. nud.
01 Genus †Aguaraya Rusconi, 1954
02 Genus †Arthricocephalites Lu, Zhang et al., 1974
03 Genus †Avalonia Walcott, 1889
04 Genus †Botomella Suvorova, 1958
05 Genus †Chakasskia Poletayeva, 1936
06 Genus †Chengkouia Lu, Chang et al., 1974
07 Genus †Feilongshania Chien & Lin, 1978
08 Genus †Gelasene Palmer, 1968
09 Genus †Giordanella Bornemann, 1891
10 Genus †Goldfieldia Palmer, 1964
11 Genus †Jangudaspis Ogienko, 1956
12 Genus †Judaiella Lermontova, 1951
13 Genus †Keeleaspis Fritz, 1972
14 Genus †Laticephalus Pokrovskaya, 1959
15 Genus †Meneghinella Boernemann, 1891
16 Genus †Metisaspina Sivov, 1955
17 Genus †Miranella Pokrovskaya, 1960
18 Genus †Nehanniaspis Fritz, 1972
19 Genus †Otekmaspis
20 Genus †Paragraulos Lu, 1941
21 Genus †Polliaxis Palmer, 1968
22 Genus †Sailycaspis Repina, 1965
23 Genus †Sekwiaspis Fritz, 1972
24 Genus †Tuyunaspis
25 Genus †Variopelta Fritz, 1972
26 Genus †Xiuqiella Lu Chang, Chien, Chu, Lin, Chou, Chien, Zhang & Wu, 1974
27 Genus †Yukonides Fritz, 1972
28 Family †Acrocephalitidae Hupé, 1953, incertae sedis
29 Family †Aldonaiidae Hupé, 1953, incertae sedis
30 Family †Amgaspididae Černyševa, 1960, incertae sedis
31 Family †Anomocarellidae Hupé, 1955, incertae sedis
32 Family †Anomocaridae Poulsen, 1927, incertae sedis
33 Family †Antagmidae Hupé, 1953, incertae sedis
34 Family †Atopidae Hupé, 1953, incertae sedis
35 Family †Auritamidae Öpik, 1967, incertae sedis
36 Family †Avoninidae Lochman, 1936, incertae sedis
37 Family †Bolaspididae Howell, 1959, incertae sedis
38 Family †Catillicephalidae Raymond, 1938, incertae sedis
39 Family †Changshaniidae Kobayashi, 1935, incertae sedis
40 Family †Cheilocephalidae Shaw, 1956, incertae sedis
41 Family †Conocoryphidae Angelin, 1854, incertae sedis
42 Family †Damesellidae Kobayashi, 1935, incertae sedis
43 Family †Diceratocephalidae Lu, 1954, incertae sedis
44 Family †Dikelocephalidae Miller, 1889, incertae sedis
45 Family †Ellipsocephaloididae Hupé, 1955, incertae sedis
46 Family †Elviniidae Kobayashi, 1935, incertae sedis
47 Family †Eurekiidae Hupé, 1953, incertae sedis
48 Family †Harpididae Whittington, 1950, incertae sedis
49 Family †Holocephalinidae Hupé, 1953, incertae sedis
50 Family †Hungaiidae Raymond, 1924, incertae sedis
51 Family †Ignotogregatidae Zhang & Jell, 1987, incertae sedis
52 Family †Inouyiidae Zhang, 1963, incertae sedis
53 Family †Isocolidae Angelin, 1854, incertae sedis
54 Family †Ityophoridae Warburg, 1925, incertae sedis
55 Family †Jamrogiidae Bentley, Jago & Cooper, 2009, incertae sedis
56 Family †Kingstoniidae Kobayashi, 1933, incertae sedis
57 Family †Lisaniidae Zhang, 1963, incertae sedis
58 Family †Llanoaspididae Lochman, 1944, incertae sedis
59 Family †Lonchocephalidae Hupé, 1953, incertae sedis
60 Family †Lorenzellidae Zhang, 1963, incertae sedis
61 Family †Mapaniidae Zhang, 1963, incertae sedis
62 Family †Menomoniidae Walcott, 1916, incertae sedis
63 Family †Missisquoiidae Hupé, 1953, incertae sedis
64 Family †Monkaspididae Kobayashi, 1935, incertae sedis
65 Family †Namanoiidae Lermontova, 1951, incertae sedis
66 Family †Nepeidae Whitehouse, 1939, incertae sedis
67 Family †Norwoodiidae Walcott, 1916, incertae sedis
68 Family †Onchonotopsidae Shaw, 1966, incertae sedis
69 Family †Papyriaspididae Whitehouse, 1939, incertae sedis
70 Family †Phylacteridae Ludvigsen & Westrop, 1989, incertae sedis
71 Family †Plethopeltidae Raymond, 1925, incertae sedis
72 Family †Polycyrtaspididae Öpik, 1967, incertae sedis
73 Family †Proasaphiscidae Zhang, 1963, incertae sedis
74 Family †Ptychaspididae Raymond, 1924, incertae sedis
75 Family †Ptychopariidae Matthew, 1887, incertae sedis [polyphyletic]
76 Family †Raymondinidae Clark, 1924, incertae sedis
77 Family †Rhyssometopidae Öpik, 1967, incertae sedis
78 Family †Sarkiidae Hupé, 1953, incertae sedis
79 Family †Shumardiidae Lake, 1907, incertae sedis
80 Family †Sunaspididae Zhang & Jell, 1987, incertae sedis
81 Family †Utiidae Kobayashi, 1935, incertae sedis
82 Family †Wuaniidae Zhang & Yuan, 1981, incertae sedis
83 Order †Eodiscida Kobayashi, 1939
84 Order †Redlichiida Richter, 1932
85 Order †Corynexochida Kobayashi, 1935
86.1 Order †Lichida Moore, 1959
86.2 Order †Odontopleurida Whittington, 1959 [monotypic]
87 Order †Phacopida Salter, 1864
88 Order †Proetida Fortey & Owens, 1975
89 Order †Aulacopleurida Adrain, 2011
90 Order †Asaphida Salter, 1864
91 Order †Olenida Adrain, 2011
92 Order †Harpida Whittington, 1959 [monotypic]