Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Hydrophiloidea Latreille, 1802
Family Hydrophilidaeᵀ Latreille, 1802 - water scavenger beetles
Genus Helophorus Fabricius, 1775
Subgenus Helophorus (Helophorus)ᵀ Leach
Helophorus (Helophorus) affinis auct.
Helophorus (Helophorus) asperatus Rey, 1885
Helophorus (Helophorus) crenatus Rey
Helophorus (Helophorus) croaticus Kuwert
Helophorus (Helophorus) discrepans Rey
Helophorus (Helophorus) dorsalis Marsham, 1802
Helophorus (Helophorus) erichsoni Bach
Helophorus (Helophorus) fallax Kuw.
Helophorus (Helophorus) flavipes (Fabricius, 1792)
Helophorus (Helophorus) fulgidicollis Motschulsky, 1860
Helophorus (Helophorus) granularis (Linnaeus, 1761)
Helophorus (Helophorus) griseus Herbst, 1793
Helophorus (Helophorus) iteratus Sharp
Helophorus (Helophorus) laticollis Thomson, 1853
Helophorus (Helophorus) longitarsis Wollaston, 1864
Helophorus (Helophorus) minutus (Fabricius, 1775)
Helophorus (Helophorus) nanus Sturm
Helophorus (Helophorus) obscurus Mulsant, 1844
Helophorus (Helophorus) pumilio auct.
Helophorus (Helophorus) pumilio Erichson
Helophorus (Helophorus) quadrisignatus Bach
Helophorus (Helophorus) redtenbacheri Kuw.
Helophorus (Helophorus) semifulgens auct., d'Orchymont
Helophorus (Helophorus) semifulgens Rey
Helophorus (Helophorus) strigifrons Thomson, 1868
Helophorus (Helophorus) villosus Duft.
Helophorus (Helophorus) walkeri Sharp
Helophorus (Helophorus) zoppae Ganglb.