Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Hydrophiloidea Latreille, 1802
Family Histeridae Gyllenhal, 1808
Genus Aeletes Horn, 1873
Genus Anapleus Horn, 1873
Genus Aphelosternus Wenzel, 1962
Genus Baconia Lewis, 1885
Genus Caerosternus LeConte, 1852
Genus Chelyoxenus Hubbard, 1894
Genus Eremosaprinus Ross, 1939
Genus Euspilotus Lewis, 1907
Genus Geocolus Wenzel, 1944
Genus Iliotona Carnochan, 1917
Genus Margarinotus Marseul, 1853
Genus Merohister Reitter, 1909
Genus Mroczkowskiella Mazur, 1984
Genus Neopachylopus Reichardt, 1926
Genus Omalodes Erichson, 1834
Genus Operclipygus Marseul, 1870
Genus "Oxysternus" Dejean, 1833, nom. nud.
Genus Peploglyptus LeConte, 1880
Genus Phelister Marseul, 1853
Genus Pseudepierus Casey, 1916
Genus Psiloscelis Marseul, 1853
Genus Reninus Lewis, 1889
Genus Spilodiscus Lewis, 1906
Genus Stictostix Marseul, 1870
Genus Terapus Marseul, 1862
Genus Teretriosoma Horn, 1873
Genus Tribalister Horn, 1873
Genus Ulkeus Horn, 1885
Genus Xerosaprinus Wenzel, 1962
Genus Xestipyge Marseul, 1862
Genus Yarmister Wenzel, 1939
1 Subfamily Niponiinae Fowler, 1912
2 Subfamily Trypaneinae
3 Subfamily Abraeinae MacLeay, 1819
4 Subfamily Saprininae Blanchard, 1845
5 Subfamily Dendrophilinae Reitter, 1909
6 Subfamily Histerinaeᵀ Gyllenhal, 1808
7 Subfamily Hetaeriinae Marseul, 1857
8 Subfamily Chlamydopsinae Bickhardt, 1914