Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Staphyliniformia Latreille, 1802
Superfamily Staphylinoideaᵀ Latreille, 1802
Family Ptiliidae Erichson, 1845
Genus Acrotrichis Motschoulsky, 1848
Genus Actidium Matthews, 1868
Genus Actinopteryx Matthews, 1872
Genus Astatopteryx Perris, 1862
Genus Baeocrara Thomson, 1859
Genus Bambara Vuillet, 1911
Genus Dilinium Casey, 1924
Genus Eurgyne Dybas, 1966
Genus Euryptilium Matth.
Genus Hydnosella Dybas, 1961
Genus Limulodes Matthews, 1867
Genus Macdonaldium Abdullah & Abdullah, 1967
Genus Micridium Motschulsky, 1869
Genus Microptilium Matth.
Genus Micrus Motschoulsky, 1848
Genus Motschulskium Matthews, 1872
Genus Nanoptilium Flach, 1889
Genus Nanosella Motschulsky, 1868
Genus Nephanes Thomson, 1859
Genus Nossidium Erichson, 1845
Genus Oligella Motschulsky
Genus Paralimulodes Bruch, 1919
Genus Plitium Bes.
Genus Porophila Dybas, 1956
Genus Ptenidium Erichson, 1845
Genus Pteryx Matthews, 1858
Genus Ptiliodina Casey, 1924
Genus Ptiliola Haldeman, 1849
Genus Ptiliolum Flach ex Seidlitz, 1888
Genus Ptiliopycna Casey, 1924
Genus Ptinella Motschulsky, 1872
Genus Ptinellodes Matthews, 1872
Genus Smicrus Matthews, 1872
Genus Suturina Dybas, 1980
Genus Throscoptilium Barber, 1924
1 Subfamily Ptiliinaeᵀ Erichson, 1845
2 Subfamily Cephaloplectinae