Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Subphylum Chelicerata Heymons, 1901
Euchelicerata Weygoldt & Paulus, 1979
Prosomapoda Lamsdell, 2013
Planaterga Lamsdell, 2013
Dekatriata Lamsdell, 2013
Sclerophorata Kamenz, Staude & Dunlop, 2011
Class Arachnida Lamarck, 1801
Micrura Hansen & Sørensen, 1904
Acaromorpha Dubinin, 1957
Subclass Acari Leach, 1817 - mites and ticks
Superorder Acariformes Zakhvatkin, 1952 - mitelike mites
Order Sarcoptiformes Reuter, 1909
Suborder Oribatida van der Hammen, 1968 - oribatid mites
Infraorder Desmonomata Woolley, 1973
Hyporder Astigmata Canestrini, 1891
Superfamily Pterolichoidea Trouessart & Mégnin, 1884
Family Pterolichidaeᵀ Trouessart & Mégnin, 1884
Subfamily Pterolichinaeᵀ Trouessart & Mégnin, 1884
01 Genus Aegothelichus Atyeo, 1979
02 Genus Afrolichus Gaud & Atyeo, 1996
03 Genus Ambodrilus Gaud & Atyeo, 1996
04 Genus Anapodema Gaud & Mouchet, 1959
05 Genus Aniacarus Gaud & Atyeo, 1990
06 Genus Aniibius Gaud & Atyeo, 1990
06.0 [Genus "Aniibus" Gaud & Atyeo, 1990]
see Genus Aniibius Gaud & Atyeo, 1990
07 Genus Anoplotarsus Gaud & Atyeo, 1996
08 Genus Apatelacarus Atyeo & Gaud, 1980
09 Genus Apexolichus Gaud & Atyeo, 1996
10 Genus Aralichus Gaud, 1966
11 Genus Aramobius Gaud & Atyeo, 1996
12 Genus Arhytidelasma Gaud & Atyeo, 1996
13 Genus Ascetolichus Pérez & Atyeo, 1990
14 Genus Aterolichus Gaud, 1980
15 Genus Atopolichus Gaud, 1980
16 Genus Atrichotibia Atyeo & Gaud, 1984
17 Genus Avenzoariurus Atyeo, 1985
18 Genus Botryaspis Atyeo, 1992
18.0 [Genus "Botryaspsis" Atyeo, 1992]
see Genus Botryaspis Atyeo, 1992
19 Genus Calvolichus Gaud & Atyeo, 1996
20 Genus Ceratothrix Trouessart, 1916
21 Genus Chelomatolichus Gaud & Atyeo, 1996
22 Genus Clastonotus Gaud, 1966
23 Genus Coccylichus Gaud, 1966
24 Genus Colinolichus Gaud & Atyeo, 1996
25 Genus Contolichus Atyeo & Gaud, 1992
26 Genus Coracopsobius Gaud & Atyeo, 1996
27 Genus Ctenodiscopus Gaud, 1980
28 Genus Cycloprotarsus Atyeo, 1992
29 Genus Cyrtonyxobius Gaud & Atyeo, 1996
30 Genus Delphinurus Gaud & Atyeo, 1996
31 Genus Diodochaetus Gaud & Atyeo, 1996
31.0 [Genus Dicranogonus Atyeo & Gaud, 1987, nom. illeg.]
see Genus Diodochaetus Gaud & Atyeo, 1996
32 Genus Distigmesikya Atyeo, Gaud & Pérez, 1984
33 Genus Doleracarus Atyeo & Gaud, 1980
34 Genus Echinofemur Atyeo & Pérez, 1984
35 Genus Echinozonus Atyeo & Pérez, 1991
36 Genus Epopolichus Gaud, 1981
37 Genus Eurypterolichus Atyeo, 1992
38 Genus Falcolichus Gaud & Atyeo, 1996
39 Genus Gabuciniella Gaud & Atyeo, 1996
40 Genus Genoprotolichus Gaud & Atyeo, 1996
41 Genus Geranolichus Gaud, 1968
42 Genus Goniodurus Atyeo, 1992
43 Genus Grallobia Hull, 1932
43.0 [Tribe Grallobiini Dubinin, 1953]
see Subfamily Pterolichinae Trouessart & Mégnin, 1884
44 Genus Grallolichus Gaud, 1960
45 Genus Gruolichus Atyeo & Windingstad, 1979
46 Genus Gymnolichus Gaud & Mouchet, 1961
47 Genus Haptepigynus Atyeo, 1992
48 Genus Heliaspis Atyeo, 1992
49 Genus Ibidonemoma Gaud & Atyeo, 1996
50 Genus Leipobius Atyeo, 1992
51 Genus Lopharalichus Gaud & Atyeo, 1996
52 Genus Lorilichus Atyeo & Gaud, 1991
53 Genus Madagascaracarus Dubinin, 1956
54 Genus Maleolichus Atyeo, 1992
55 Genus Mastigodiscus Atyeo & Gaud, 1984
56 Genus Mayracarus Atyeo, 1992
57 Genus Megapodobius Atyeo, 1992
58 Genus Mesolichus Trouessart, 1899
59 Genus Musophagobius Gaud & Atyeo, 1996
60 Genus Nyctibiolichus Atyeo, 1979
61 Genus Opisthocomacarus Dubinin, 1955
61.0 [Genus "Opisthocomacarus" Dubinin, 1953]
see Genus Opisthocomacarus Dubinin, 1955
62 Genus Ornatumerus Gaud & Atyeo, 1996
63 Genus Oxygynurus Atyeo, 1992
64 Genus Pararalichus Atyeo, 1989
65 Genus Pavolichus Gaud, 1965
66 Genus Pereziella Atyeo, 1990
67 Genus Periexocaulus Gaud & Atyeo, 1996
68 Genus Phasidolichus Gaud & Atyeo, 1996
69 Genus Phycoferus Atyeo & Pérez, 1991
70 Genus Prionoturus Atyeo, 1992
71 Genus Protolichus Trouessart, 1884
72 Genus Pseudalloptes Mégnin & Trouessart, 1883
72.0 [Genus Cynthiura Hull, 1932]
see Genus Pseudalloptes Mégnin & Trouessart, 1883
72.0 [Tribe Pseudalloptini Dubinin, 1953]
see Subfamily Pterolichinae Trouessart & Mégnin, 1884
73 Genus Pseudalloptinus Dubinin, 1956
74 Genus Pseudolichus Atyeo & Gaud, 1992
75 Genus Psittocolus Gaud & Atyeo, 1996
76 Genus Pterolichusᵀ Robin, 1868
77 Genus Pterygocrusolichus Dubinin, 1955
78 Genus Ptyssalgopus Gaud & Atyeo, 1996
79 Genus Rhytidelasma Gaud, 1966
80 Genus Scolaralichus Gaud & Atyeo, 1996
81 Genus Scopusacarus Dubinin, 1956
81.0 [Genus "Scopusacarus" Dubinin, 1953]
see Genus Scopusacarus Dubinin, 1956
81.0 [Genus Scopusacarus Gaud & Mouchet, 1959]
see Genus Scopusacarus Dubinin, 1956
82 Genus Sideroferus Gaud & Atyeo, 1996
83 Genus Stakyonemus Atyeo & Gaud, 1971
84 Genus Struthiopterolichus Dubinin, 1955
84.0 [Genus "Struthiopterolichus" Dubinin, 1953]
see Genus Struthiopterolichus Dubinin, 1955
85 Genus Synapsilobus Gaud & Atyeo, 1996
86 Genus Talegallobius Atyeo, 1992
87 Genus Tamnurolichus Gaud & Atyeo, 1996
88 Genus Tanyaralichus Gaud & Atyeo, 1996
89 Genus Tanysomacarus Atyeo, 1992
90 Genus Tetraolichus Atyeo & Gaud, 1992
91 Genus Titanolichus Gaud & Atyeo, 1996
92 Genus Touracobia Gaud & Mouchet, 1959
93 Genus Turnixacarus Gaud & Atyeo, 1996
94 Genus Uropsittacolichus Atyeo & Pérez, 1982
95 Genus Xoloptes Canestrini, 1879
x [Genus "Ciconiacarus" Dubinin, 1953, nom. nud.]
see Genus Ciconiacarus Dubinin, 1956