Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade "neomura" Cavalier-Smith 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
Clade neozoa Cavalier-Smith 1993
Clade scotokaryotes Cavalier-Smith 2000
Clade podiates Cavalier-Smith 2012
[crown] Clade Amorphea Adl et al. 2012
[crown] Clade Opisthokonta Cavalier-Smith 1987
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Geotrupidae Latreille, 1802
Genus Bolbocerosoma Schaeffer, 1906
Genus Ceratophyus Fischer de Waldheim, 1824
Genus Mycotrupes LeConte, 1866
Genus Peltotrupes Blanchard, 1888
1 Subfamily Taurocerastinae Germain, 1897
2 Subfamily Bolboceratinae Mulsant, 1842
3 Subfamily Geotrupinaeᵀ Latreille, 1802