Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Superfamily Tortricoidea Latreille, 1802, sedis mutabilis [monotypic]
Family Tortricidaeᵀ Latreille, 1802 - leafroller moths
Subfamily Olethreutinae Walsingham, 1895
Genus Acanthoclita Diakonoff, 1982
Genus Acroclita Lederer, 1859
Genus Adenoneura Walsingham, 1907
Genus Aemulatrix Diakonoff, 1982
Genus Age Diakonoff, 1982
Genus Agriophanes Meyrick, 1930
Genus Ahmosia Heinrich, 1926
Genus Alcina Clarke, 1976
Genus Alischirnevaia Kocak, 1981
Genus Allodapella Diakonoff, 1948
Genus Allohermenias Diakonoff, 1953
Genus Anathamna Meyrick, 1911
Genus Ancylis Hübner, [1825]
Genus Andrioplecta Obraztsov, 1968
Genus Anthozela Meyrick, 1913
Genus Antichlidas Meyrick, 1931
Genus Antictenista Meyrick, 1927
Genus Apolychrosis Amsel, 1962
Genus Argyroploce Hübner, [1825]
Genus Argyroptocha Diakonoff, 1967
Genus Articolla Meyrick, 1907
Genus Artiphanes Diakonoff, 1966
Genus Asketria Fal'kovich, 1964
Genus Assulella Kuznetzov, 1973
Genus Astenodes Kuznetsov, 1966
Genus Astronauta Diakonoff, 1983
Genus Atrypsiastis Meyrick, 1932
Genus Bactrostoma Diakonoff, 1960
Genus Bakia Kocak, 1981
Genus Barbara Heinrich, 1923
Genus Basigonia Diakonoff, 1983
Genus Bathrotoma Meyrick, 1881
Genus Bipartivalva Kuznetzov, 1988
Genus Blastesthia Obraztsov, 1960
Genus Blastopetrova Liu & Wu, 1987
Genus Brachiocera Diakonoff, 1959
Genus Brachioxena Diakonoff, 1968
Genus Bucephalacra Diakonoff, 1970
Genus Cacocharis Walsingham, 1892
Genus Camptrodoxa Meyrick, 1925
Genus Catastega Clemens, 1861
Genus Cellifera Diakonoff, 1967
Genus Centroxena Diakonoff, 1971
Genus Chimoptesis Powell, 1964
Genus Cimeliomorpha Diakonoff, 1966
Genus Cirrilaspeyresia Razowski, 1961
Genus Cirriphora Obraztsov, 1951
Genus Clavigesta Obraztsov, 1946
Genus Coccothera Meyrick, 1914
Genus Coccyx Treitschke, 1829
Genus Coenobiodes Kuznetzov, 1973
Genus Collicularia Obraztsov, 1960
Genus Collogenes Meyrick, 1931
Genus Coniostola Diakonoff, 1961
Genus Corticivora Clarke, 1951
Genus Cosmetra Diakonoff, 1977
Genus Cosmorrhyncha Meyrick, 1913
Genus Crimnologa Meyrick, 1920
Genus Crocidosema Zeller, 1847
Genus Crocostola Diakonoff, 1953
Genus Crusimetra Meyrick, 1912
Genus Cryptaspasma Walsingham, 1900
Genus Cryptophlebia Walsingham, 1900[1899]
Genus Cydia Hübner, [1825]
Genus Cyphophanes Meyrick, 1937
Genus Dasodis Diakonoff, 1982
Genus Dasybregma Diakonoff, 1983
Genus Demeijerella Diakonoff, 1954
Genus Dichrorampha Guenee, 1845
Genus Dicnecidia Diakonoff, 1982
Genus Dierlia Diakonoff, 1976
Genus Dinogenes Meyrick, 1934
Genus Diplonearcha Meyrick, 1914
Genus Diplosemaphora Diakonoff, 1982
Genus Dolichohedya Diakonoff, 1970
Genus Doliochastis Meyrick, 1920
Genus Dracontogena Diakonoff, 1970
Genus Duessa Clarke, 1976
Genus Eccopsis Zeller, 1852
Genus Eccoptocera Walsingham, 1907
Genus Ecdytolopha Zeller, 1875
Genus Electresia Kuznetzov, 1941
Genus Embolostoma Diakonoff, 1977
Genus Emrahia Kocak, 1981
Tribe Enarmoniini
Genus Enarmoniodes Ghesquière, 1940
Genus Enveryucelia Kocak, 1981
Genus Epibactra Ragonot, 1894
Genus Epiblema Hübner, 1825
Genus Epinotia Hübner, 1825
Genus Episimoides Diakonoff, 1957
Genus Episimus Walsingham, 1892
Genus Eriopsela Guenée, 1845
Genus Eucoenogenes Meyrick, 1939
Genus Eucosma Hübner, 1823
Genus Eucosmogastra Diakonoff, 1975
Genus Eucosmoides Obraztsov, 1946
Genus Eucosmomorpha Obraztsov, 1951
Genus Eucosmophyes Diakonoff, 1982
Genus Eumarozia Heinrich, 1926
Genus Evora Heinrich, 1926
Genus Exartema Clemens, 1860
Genus Foveifera Obraztsov, 1946
Genus Fulcrifera Danilevskii & Kuznetsov, 1968
Genus Ganabalia Diakonoff, 1975
Genus Gibberifera Obraztsov, 1946
Genus Goditha Heinrich, 1926
Genus Grapholita Treitschke, 1829
Tribe Grapholitini Guenee, 1845
Genus Gravitarmata Obraztsov, 1946
Genus Gretchena Heinrich, 1923
Genus Gypsonoma Meyrick, 1895
Genus Gypsonomoides Obraztsov, 1946
Genus Heleanna Clarke, 1976
Genus Helictophanes Meyrick, 1882
Genus Hendecaneura Walsingham, 1900
Genus Hendecasticha Meyrick, 1882
Genus Hermenias Meyrick, 1911
Genus Herpystis Meyrick, 1911
Genus Hikagehamakia Oku, 1974
Genus Hilaroptila Diakonoff, 1970
Genus Hiroshiinoueana Kawabe, 1978
Genus Hopliteccopsis Diakonoff, 1963
Genus Hylotropha Turner, 1946
Genus Hystrichophora Walsingham, 1879
Genus Hystrichoscelus Walsingham, 1900
Genus Icelita Bradley, 1957
Genus Jerapowellia Miller, 1995
Genus Karacaoglania Kocak, 1981
Genus Kennelia Rebel, 1901
Genus Larisa Miller, 1979
Genus Lathronympha Meyrick, 1926
Genus Leguminivora Obraztsov, 1960
Genus Lepidunca Diakonoff, 1984
Genus Lepteucosma Diakonoff, 1971
Genus Licigena Diakonoff, 1982
Genus Loboschiza Diakonoff, 1967
Genus Lorita Busck, 1939
Genus Macraesthetica Meyrick, 1932
Genus Megaherpystis Diakonoff, 1969
Genus Mehteria Kocak, 1981
Genus Melanalopha Diakonoff, 1941
Genus Mesocallyntera Diakonoff, 1959
Genus Metacosma Kuznetsov, 1985
Genus Metaselena Diakonoff, 1939
Genus Mevlanaia Kocak, 1981
Tribe Microcorsini
Genus Microsarotis Diakonoff, 1982
Genus Mimarsinania Kocak, 1981
Genus Mystogenes Meyrick, 1930
Genus Nenomoshia Clarke, 1976
Genus Neoanathamna Kawabe, 1978
Genus Neohermenias Diakonoff, 1966
Genus Neorrhyncha Aarvik, 2004
Genus Neostatherotis Oku, 1974
Genus Niphadophylax Diakonoff, 1992
Genus Noduliferola Kuznetzov, 1973
Genus Notocelia Hübner, 1825
Genus Nuntiella Kuznetzov, 1971
Genus Omiostola Meyrick, 1922
Genus Oriodryas Turner, 1925
Genus Osthelderiella Obraztsov, 1961
Genus Pammene Hübner, 1825
Genus Pammenemima Diakonoff, 1982
Genus Pammenodes Danilevskii & Kuznetsov, 1968
Genus Parachanda Meyrick, 1927
Genus Paralobesia Obraztsov, 1953
Genus Parapammene Obraztsov, 1960
Genus Parepisimia Diakonoff, 1975
Genus Pelochrista Lederer, 1859
Genus Penestostoma Diakonoff, 1992
Genus Penthostola Diakonoff, 1978
Genus Peridaedala Meyrick, 1925
Genus Periphoeba Bradley, 1957
Genus Phalarocarpa Meyrick, 1937
Genus "Phaneta" Stephens, 1852, nom. nud.
Genus Plutographa Diakonoff, 1970
Genus Potiosa Diakonoff, 1965
Genus Procoronis Meyrick, 1911
Genus Protancylis Diakonoff, 1983
Genus Proteoteras Riley, 1882
Genus Protobactra Diakonoff, 1964
Genus Psegmatica Meyrick, 1930
Genus Pseudexentera Heinrich, 1940
Genus Pseudoclita Bradley, 1957
Genus Pseudococcyx Swatschek, 1958
Genus Pseudogalleria Ragonot, 1884
Genus Pseudophiaris Obraztsov, 1961
Genus Pternidora Meyrick, 1911
Genus Retinia Guenée, 1845
Genus Rhectogonia Diakonoff, 1966
Genus Rhodotoxotis Diakonoff, 1992
Genus Rhopalovalva Kuznetsov, 1964
Genus Rhopobota Lederer, 1859
Genus Rhyacionia Hübner, 1825
Genus Riculoides Pastrana, 1952
Genus Ruthita Clarke, 1976
Genus Salsolicola Kuznetsov, 1960
Genus Satronia Heinrich, 1926
Genus Selania Stephens, 1834
Genus Semnostola Diakonoff, 1959
Genus Semutophila Tuck, 1986
Genus Sereda Heinrich, 1923
Genus Sillybiphora Kuznetsov, 1964
Genus Sonia Heinrich, 1923
Genus Spanistoneura Diakonoff, 1982
Genus "Spilonota" Stephens, 1829, nom. nud.
Genus Stenentoma Diakonoff, 1969
Genus Strepsicrates Meyrick, 1888
Genus Strophedra Herrich-Schäffer, [1854]
Genus Strophedromorpha Diakonoff, 1976
Genus Stygitropha Diakonoff, 1983
Genus Suleima Heinrich, 1923
Genus Syropetrova Diakonoff, 1970
Genus Tetramoera Diakonoff, 1967
Genus Thaumatoptyla Diakonoff, 1984
Genus Thiodia Hübner, [1825]
Genus Thiodiodes Obraztsov, 1964
Genus Thylacandra Diakonoff, 1963
Genus Thymioptila Diakonoff, 1971
Genus Tia Heinrich, 1926
Genus Titanotoca Diakonoff, 1984
Genus Tokuana Kawabe, 1978
Genus Trachyschistis Meyrick, 1921
Genus Tritopterna Meyrick, 1921
Genus Tsinilla Heinrich, 1931
Genus Ukamenia Oku, 1981
Genus Xenopotamia Diakonoff, 1983
Genus Yunusemreia Kocak, 1981
Genus Zeiraphera Treitschke, 1829
Genus Zomariana Diakonoff, 1984
1.1 Tribe Gatesclarkeanini
1.2 Tribe Bactrini Falkovitch, 1962c
1.3 Tribe Endotheniini
1.4 Tribe Olethreutiniᵀ Obraztsov, 1946
2 Tribe Eucosmini
3 Tribe Laspeyresiini