Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade dorsates Cavalier-Smith, 2021
                   Clade podiates Cavalier-Smith, 2012
                     [crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                       [crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                         [crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                       Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                             Tactopoda Budd, 2001
                                               Phylum Arthropoda Siebold, 1848 - arthropods
                                                 Euarthropoda
                                                   Antennulata
                                                     Subphylum Mandibulata Snodgrass, 1938
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                         Altocrustacea
                                                           Miracrustacea
                                                             Superclass Hexapoda Latreille, 1825
                                                               Cercophora
                                                                 Class Insecta Linnaeus, 1758 - insects
                                                                   Subclass Dicondylia Haeckel, 1866
                                                                     Infraclass Pterygota Lang, 1888
                                                                       Metapterygota
                                                                         Neoptera Martynov, 1923
                                                                           Eumetabola
                                                                             Holometabola
                                                                               Order Coleoptera Linnaeus, 1758 - beetles
                                                                                 Suborder Polyphaga Emery, 1886
                                                                                   Series Scarabaeiformia Crowson, 1960
                                                                                     Superfamily Scarabaeoideaᵀ Latreille, 1802
                                                                                       Family Geotrupidae Latreille, 1802
                                                                                         Subfamily Bolboceratinae Mulsant, 1842
                                                                                           Genus Bolbocerastes Cartwright, 1953
                                                                                           Genus Bolborhombus Cartwright, 1953
                                                                                           Genus Bradycinetulus Cockerell, 1906
                                                                                           Genus Elephastomus MacLeay, 1819
                                                                                           1 Tribe Bolboceratiniᵀ Mulsant, 1842
                                                                                           2 Tribe Athyreini Howden & Martínez, 1963
                                                                                           3 Tribe Bolbochromini Nikolajev, 1970
                                                                                           4 Tribe Gilletinini Nikolajev, 1990
                                                                                           5 Tribe Eubolbitini Nikolajev, 1970
                                                                                           6 Tribe Bolbelasmini Nikolajev, 1996
                                                                                           7 Tribe Australobolbini Nikolajev, 1996
                                                                                           8 Tribe Eucanthini Nikolajev, 2002b
                                                                                           9 Tribe Stenaspidiini Nikolajev, 2003