Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Cetoniinae Leach, 1815
Genus Anatropis Casey, 1915
Genus Cotinis Burmeister, 1842
Genus Epicometis Burmeister, 1842
Genus Genuchinus Westwood, 1874
Genus Gnorimella Casey, 1915
Genus Lissomelas Bates, 1889
Genus Psilocnemis Burmeister, 1842
Genus Stephanucha Burmeister, 1842
Genus Trichiotinus Casey, 1915
Genus Trigonopeltastes Burmeister, 1840
01 Tribe Valgini Mulsant, 1842
02 Tribe Trichiini Fleming, 1821
03 Tribe Cremastocheilini Burmeister & Schaum, 1841
04 Tribe Xiphoscelidini Burmeister, 1842
05 Tribe Stenotarsiini Kraatz, 1880
06 Tribe Schizorhinini Burmeister, 1842
07 Tribe Goliathini Griffith & Pidgeon, 1832
08 Tribe Cetoniiniᵀ Leach, 1815
09 Tribe Gymnetini Kirby, 1827
10 Tribe Diplognathini Burmeister, 1842
11 Tribe Phaedimini Schoch, 1894
12 Tribe Taenioderini Miksic, 1976