Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Genus Aganhyboma Kolbe, 1893
Genus Amphiceratodon Huchet, 2000
Genus Anomalacra Casey, 1915
Genus Arachnodes Westwood, 1843
Genus Boreocanthon Halffter, 1958
?Genus Catharsius Hope, 1837
Genus Chalcosoma Hope, 1837
Genus Chaunocolus Saylor, 1937
Genus Chlorixantha Bates, 1889
Genus Chnaunanthus Burmeister, 1844
Genus Chrysina Kirby, 1828
Genus Cineretis Schürhoff, 1937
Genus Cotalpa Burmeister, 1844
Genus Diastictus Mulsant, 1842
Genus Dinacoma Casey, 1890
Genus Fossocarus Howden, 1967
Genus Gnorimus Lepeletier & Serville, 1828
Genus Gronocarus Schaeffer, 1927
Genus Hypothyce Hardy, 1974
Genus Hypotrichia LeConte, 1862
Genus Ignambia Heller, 1916
Genus Liocola Thomson, 1859
Genus Maladera Mulsant, 1871
Genus Megathopa Eschscholtz, 1822
Genus Miltotrogus Reitter, 1902
Genus Mimela Kirby, 1823
Genus Nefoncerus Saylor, 1938
Genus †Oryctoantiquus Ratcliffe & D. Smith, 2005
Genus Oxythyrea Mulsant, 1842
Genus Parabyrsopolis Ohaus, 1915
Genus Paracotalpa Ohaus, 1915
Genus Parathyce Hardy, 1974
Genus Phobetus LeConte, 1856
Genus Phyllophaga Harris, 1827
Genus Plectrodes Horn, 1867
Genus Pleurophorus Mulsant, 1842
Genus Potosia Mulsant & Rey, 1871
Genus Prodontria Broun, 1904
Genus Protaetia Burmeister, 1842
Genus Pseudocotalpa Hardy, 1971
Genus Strigoderma Burmeister, 1844
Genus Thyce LeConte, 1856
Genus Tropinota Mulsant, 1842
01 Subfamily †Lithoscarabaeinae Nikolajev, 1992
02 Subfamily Chironinae Blanchard, 1845
03.1 Subfamily Aegialiinae Laporte, 1840
03.2 Subfamily Eremazinae Iablokoff-Khnzorian, 1977
04 Subfamily Aphodiinae Leach, 1815
05 Subfamily Aulonocneminae Janssens, 1946
06 Subfamily Termitotroginae Wasmann, 1918
07 Subfamily Scarabaeinaeᵀ Latreille, 1802
08 Subfamily †Prototroginae Nikolajev, 2000
09 Subfamily †Cretoscarabaeinae Nikolajev, 1995
10 Subfamily Dynamopodinae Arrow, 1911, nom. emend.
11 Subfamily Phaenomeridinae Erichson, 1847, nom. emend.
12 Subfamily Orphninae Erichson, 1847
13 Subfamily Allidiostomatinae Arrow, 1940, nom. emend.
14 Subfamily Aclopinae Blanchard, 1850
15 Subfamily Melolonthinae Leach, in Samouelle, 1819
16 Subfamily Rutelinae MacLeay, 1819
17 Subfamily Dynastinae MacLeay, 1819
18 Subfamily Cetoniinae Leach, 1815