Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Melolonthinae Leach, in Samouelle, 1819
Genus Amphimallon Berthold, 1827
?Genus Anoxia Laporte, 1833
Genus Popilia Leach, 1829
01 Genus Diaphylla Erichson, 1847
02 Genus Hilarianus Blanchard
03 Genus Manonychus Moser, 1919
04 Genus Pseudoisonychus Frey
05 Tribe Pachypodini Erichson, 1840
06 Tribe Lichniini Burmeister, 1844
07 Tribe Euchirini Hope, 1840
08 Tribe Systellopini Sharp, 1877
09 Tribe Chasmatopterini Lacordaire, 1856
10 Tribe Oncerini LeConte, 1861
11 Tribe Podolasiini Howden, 1997
12 Tribe Ablaberini Blanchard, 1850
13 Tribe Sericini Kirby, 1837
14 Tribe Phyllotocidiini Britton, 1957
15 Tribe Diphucephalini Laporte, 1840
16 Tribe Comophorinini Britton, 1957
17 Tribe Stethaspini Burmeister, 1855
18 Tribe Automoliini Britton, 1978
19 Tribe Maechidiini Burmeister, 1855
20 Tribe Liparetrini Burmeister, 1855
21 Tribe Scitalini Britton, 1957
22 Tribe Pachytrichini Burmeister, 1855
23 Tribe Sericoidini Erichson, 1847
24 Tribe Heteronychini Lacordaire, 1856
25 Tribe Diplotaxini Kirby, 1837
26 Tribe Melolonthiniᵀ Leach, in Samouelle, 1819
27 Tribe Pachydemini Burmeister, 1855, nom. protect.
28.1 Tribe Macrodactylini (Kirby, 1837)
28.2 Tribe Dichelonychini (Burmeister, 1855)
29 Tribe Diphycerini Medvedev, 1952
30 Tribe Hopliini Latreille, 1829
31 Tribe Colymbomorphini Blanchard, 1850
32 Tribe Tanyproctini Erichson 1847
33 Tribe †Cretomelolonthini Nikolajev, 1998