Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Melolonthinae Leach, in Samouelle, 1819
Tribe Macrodactylini (Kirby, 1837)
01 Genus Agaocnemis Moser, 1918
02 Genus Alvarinus [Blanchard], 1850
03 Genus Ancistrosoma Curtis, 1835
04 Genus Anomonyx Saylor, 1940
05 Genus Anoplosiagum [Blanchard], 1850
06 Genus Astaenosiagum Martínez, 1957
07 Genus Barybas Blanchard, 1850
08 Genus Calodactylus [Blanchard], 1850
09 Genus Ceraspis Le Peletier & Serville, 1828
10 Genus Ceratolontha Arrow, 1948
11 Genus Chariodactylus Moser, 1919
12 Genus Chariodema [Blanchard], 1850
13 Genus Chremastodus Solier, 1851
14 Genus Clavipalpus Laporte, 1833
15 Genus Ctenotis Burmeister, 1855
16 Genus Dasyus Lepeletier & Serville, 1828
17 Genus Dicrania Lepeletier & Serville, 1828
18 Genus Gama Blanchard
19 Genus Gastrohoplus Moser, 1921
20 Genus Hercitis Burmeister, 1855
21 Genus Hieritis Burmeister, 1855
22 Genus Isonychus Mannerheim, 1829
23 Genus Issacaris Fairmaire, 1889
24 Genus Macrodactylusᵀ Dejean, 1821
25 Genus Manodactylus Moser, 1919
26 Genus Manopus Laporte, 1840
27 Genus Oedichira Burmeister, 1855
28 Genus Pectinosoma Arrow, 1913
29 Genus Plectris Le Peletier & Serville, 1828a
30 Genus Pristerophora Harold, 1869
31 Genus Rhinaspis Perty, 1830
32 Genus Schizochelus Blanchard