Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Cucujiformia Lameere, 1938
Superfamily Tenebrionoidea Latreille, 1802
Family Tenebrionidaeᵀ Latreille, 1802 - darkling beetles
Genus Aconobius Casey, 1895
Genus "Adelina" Dejean, 1835, nom. nud.
Genus Adelonia de Laporte, 1840
Genus Alaephus Horn, 1870
Genus Alaudes Horn, 1870
Genus Alegoria de Laporte, 1840
Genus Aletha Champion, 1888
Genus Alobates Motschoulsky, 1872
Genus Alphitobius Stephens, 1829
Genus Alphitophagus Stephens, 1832
Tribe Amarygmini (Gistel, 1848)
Genus Ammodonus Mulsant & Rey, 1859
Genus Anaedus Blanchard, 1845
Genus Anamphidora Casey, 1924
Genus Andrimus Casey, 1891
Genus Androchirus Leconte, 1862
Genus Anepsius LeConte, 1851
Genus Apocrypha Eschscholtz, 1831
Genus Apsida Lacordaire, 1859
Genus Araeoschizus LeConte, 1851
Genus Argoporis Horn, 1870
Genus Armalia Casey, 1907
Genus Arthromacra Kirby, 1837
Genus Asida Latreille, 1802
Genus Asidina Casey, 1912
Genus Asidopsis Casey, 1912
Genus Astrotus LeConte, 1858
Genus Auchmobius LeConte, 1851
Genus Batuliodes Casey, 1907
Genus Batuliomorpha Doyen, 1987
Genus Batulius LeConte, 1851
Genus Biomorphus Motschoulsky, 1872
Genus Bius Mulsant, 1854
Genus Blapstinus Sturm, 1826
Tribe Blaptini Leach, 1815
Tribe Bolitophagini (Kirby, 1837)
Genus Bolitophagus Illiger, 1798
Genus Bolitotherus Candèze, 1861
Genus Bothrotes Casey, 1907
Genus Branchus LeConte, 1862
Genus Calognathus Guérin, 1837
Genus Calymmus Pascoe, 1871
Genus Camaria Lepeletier & Serville, 1825
Genus Cardiosis Deyrolle, 1867
Genus Centrioptera Mannerheim, 1843
Genus Centronopus Solier, 1848
Genus Cerenopus LeConte, 1851
Genus Charisius Champion, 1888
Genus Cheirodes Gené, 1839
Genus Chilometopon Horn, 1874
Genus Choerodes Leidy, 1852
Genus Cibdelis Mannerheim, 1843
Genus Clamoris des Gozis, 1886
Genus Cnemeplatia Costa, 1847
Genus Cnemodinus Cockerell, 1906
Genus Coelocnemis Mannerheim, 1843
Genus Coelus Eschscholtz, 1829
Genus Conibiosoma Casey, 1890
Genus Conibius LeConte, 1851
Genus Coniontis Eschscholtz, 1829
Genus Conisattus Casey, 1895
Genus Conoecus Horn, 1885
Genus Corticeus Piller & Mitterpacher, 1783
Genus Craniotus LeConte, 1851
Genus Cryptadius LeConte, 1851
Genus Crypticus Latreille, 1817
Genus Cryptoglossa Solier, 1837
Genus Cteniopus Solier, 1835
Genus Cuphotes Champion, 1887
Genus Cybotus Casey, 1891
Genus Cyclosattus Casey, 1892
Genus "Cymatothes" Dejean, 1834, nom. nud.
Genus Cynaeus LeConte, 1862
Genus Cyrtomius Casey, 1907
Tribe Diaperini Latreille, 1802
Genus Dioedus Leconte, 1862
Genus Doliodesmus Spilman, 1967
Genus Edrotes LeConte, 1851
Genus Eleates Casey, 1886
Genus Eledona Latreille, 1796
Genus Eleodes Eschscholtz, 1829
Genus Eleodimorpha Blaisdell, 1909
Tribe Eleodini
Genus Embaphion Say, 1824
Genus Emmenastus Motschulsky, 1845
Genus Ephalus LeConte, 1862
Genus Epitragodes Casey, 1890
Genus Eschatomoxys Blaisdell, 1935
Genus Eschatoporis Blaisdell, 1906
Genus Eulabis Eschscholtz, 1829
Genus Eupezus
Genus Eupsophulus Cockerell, 1906
Genus Eurymetopon Eschscholtz, 1831
Genus Eusattus LeConte, 1851
Genus Eutochia LeConte, 1862
Genus Glyptasida Casey, 1912
Genus Glyptotus LeConte, 1858
Genus Gnathocerus Thunberg, 1814
Genus Gonasida Casey, 1912
Genus Gonocephalum Chevrolat, 1849
Genus Haplandrus LeConte, 1866
Tribe Helaeini
Tribe Helopini (Latreille, 1802)
Genus Helops Fabricius, 1775
Genus Hylocrinus Casey, 1907
Genus Hymenochara Campbell, 1978
Genus Hymenorus Mulsant, 1851
Genus "Hypogena" Dejean, 1834, nom. nud.
Genus Hypophloeus
Genus Idiobates Casey, 1890
Genus Iphthiminus Spilman, 1973
Genus Isomira Mulsant, 1856
Genus Knausia Fall, 1931
Genus Lariversius Blaisdell, 1947
Genus Latheticus Waterhouse, 1880
Genus Leichenum Blanchard, 1845
Genus Lepidochora Gebien, 1938
Genus Liodema Horn, 1870
Genus Lobometopon Casey, 1907
Genus Lobopoda Solier, 1835
Genus Lyphia Mulsant & Rey, 1859
Genus Lystronichus Latreille, 1829
Genus Macellocerus Solier, 1848
Genus Mecysmus Horn, 1870
Genus Megasida Casey, 1912
Genus Megeleates Casey, 1895
Genus Melanastus Casey, 1907
Genus Melanimon Steven, 1829
Genus Melasmana Strand, 1935
Genus Menoeceus Champion, 1888
Genus Meracantha Kirby, 1837
Genus Merinus LeConte, 1866
Genus Metaclisa Duval, 1861
Genus Metopoloba Casey, 1907
Genus Metoponium Casey, 1907
Genus Micromes Casey, 1907
Genus Microschatia Solier, 1836
Genus Mycetochara Berthold, 1827
Genus Mycotrogus Horn, 1870
Genus Nautes Pascoe, 1866
Genus Neatus LeConte, 1862
Genus Neobaphion Blaisdell, 1925
Genus Neomida Latreille, 1829
Genus Nocibiotes Casey, 1895
Genus Notibius LeConte, 1851
Genus Nyctoporis Eschscholtz, 1831
Tribe Nyctozoilini
Genus Oenopion Champion, 1885
Genus Onychomira Campbell, 1984
Genus Onymacris Allard, 1885
Tribe Opatrini Brullé, 1832
Genus Opatrinus Latreille, 1829
Genus Oxygonodera Casey, 1907
Genus Palorus Mulsant, 1854
Genus Parasida Casey, 1912
Genus Paratenetus Spinola, 1844
Genus Pechalius Casey, 1907
Genus Pedinus Latreille, 1796
Tribe Pentaphyllini
Genus "Pentaphyllus" Dejean, 1821, nom. nud.
Genus Phaleria Latreille, 1802
Genus Phedius Champion, 1888
Genus Phegoneus Casey, 1907
Genus Philolithus Lacordaire, 1859
Genus Philorea Erichson, 1834
Genus Phrenapates Gray, 1832
Genus Phylan
Genus Phymatestes Pascoe, 1867
Genus Platydema Laporte & Brullé, 1831
Genus Polemiotus Casey, 1907
Genus Polopinus Casey, 1924
Genus Polposipus Solier, 1848
Genus Polypleurus Eschscholtz, 1831
Genus Prateus LeConte, 1862
Genus Promethis Pascoe, 1869
Genus Prostenus Latreille, 1825
Genus Pseudephalus Casey, 1924
Genus Pycnocerus Westwood, 1841
Genus Rhipidandrus LeConte, 1862
Genus Scaphidema Redtenbacher, 1849
Genus Schizillus Horn, 1874
Genus Schoenicus LeConte, 1866
Genus Scotobaenus LeConte, 1859
Genus Sepidium Fabricius, 1775
Genus Sitophagus Mulsant, 1854
Genus Statira Serville, 1825
Genus Stenochidus LeConte, 1862
Genus Stenomax Allard, 1876
Genus Stenomorpha Solier, 1836
Genus Stenosides Solier, 1836
Genus Stenotrichus LeConte, 1862
Genus Steriphanus Casey, 1907
Genus Sternodes Fischer-Waldheim, 1837
Genus Stethasida Casey, 1912
Genus Stibia Horn, 1870
Tribe Strongyliini
Genus Strongylium Kirby, 1818
Genus Tagalus Gebien, 1914
Genus Talanus Mäklin, 1878
Genus Tanychilus Newman, 1838
Genus Tarpela Bates, 1870
Genus Tedinus Casey, 1891
Genus Telabis Casey, 1890
Genus Telesicles Champion, 1888
Genus Tharsus LeConte, 1862
Genus Threnus Motschulsky, 1870
Genus Tonibius Casey, 1895
Tribe Toxicini Oken, 1843
Genus Trachyscelis Latreille, 1809
Genus Tribolium MacLeay, 1825
Genus Trichiasida Casey, 1912
Genus Trichiotes Casey, 1907
Genus Trichoton Hope, 1840
Genus Trimytis LeConte, 1851
Genus Triorophus LeConte, 1851
Genus Triphalopsis Blaisdell, 1923
Genus Trogloderus LeConte, 1879
Genus Typhlusechus Linell, 1897
Genus Uloma Dejean, 1821
Tribe Ulomini Blanchard, 1845
Genus Ulomoides Blackburn, 1888
Genus Ulus Horn, 1870
Genus Upis Fabricius, 1792
Genus Xylopinus LeConte, 1862
Genus Zophobas Blanchard, 1845
1 Subfamily Tentyriinae
2 Subfamily Alleculinae (Laporte, 1840)
3 Subfamily Lagriinae (Latreille, 1825)
4 Subfamily Tenebrioninaeᵀ Latreille, 1802
5 Subfamily Zolodininae Watt, 1874