Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Cucujiformia Lameere, 1938
Superfamily Tenebrionoidea Latreille, 1802
01 Family Mycetophagidae Leach, 1815
02 Family Archeocrypticidae Kaszab, 1964
03 Family Pterogeniidae Crowson, 1953
04 Family Ciidae Leach, 1819
05 Family Tetratomidae Billberg, 1820
06 Family Melandryidae Leach, 1815
07 Family Mordellidae Latreille, 1802
08 Family Ripiphoridae Gemminger, 1870 (1853)
09 Family Zopheridae Solier, 1834
10 Family Ulodidae Pascoe, 1869
11 Family Promecheilidae Lacordaire, 1859
12 Family Chalcodryidae Watt, 1974
13 Family Trachelostenidae Lacordaire, 1859
14 Family Tenebrionidaeᵀ Latreille, 1802 - darkling beetles
15 Family Prostomidae Thomson, 1859
16 Family Synchroidae Lacordaire, 1859
17 Family Stenotrachelidae Thomson, 1859
18 Family Oedemeridae Latreille, 1810
19 Family Meloidae Gyllenhal, 1810 - blister beetles
20 Family Mycteridae Oken, 1843
21 Family Boridae Thomson, 1859
22 Family Trictenotomidae Blanchard, 1845
23 Family Pythidae Solier, 1834
24 Family Pyrochroidae Latreille, 1807
25 Family Salpingidae Leach, 1815
26 Family Anthicidae Latreille, 1819
27 Family Aderidae Winkler, 1927
28 Family Scraptiidae Gistel, 1848