Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 Clade neozoa Cavalier-Smith 1993
                   Clade scotokaryotes Cavalier-Smith 2000
                     Clade podiates Cavalier-Smith 2012
                       [crown] Clade Amorphea Adl et al. 2012
                         [crown] Clade Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                       Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                             Tactopoda Budd, 2001
                                               Phylum Arthropoda Siebold, 1848 - arthropods
                                                 Euarthropoda
                                                   Antennulata
                                                     Subphylum Mandibulata Snodgrass, 1938
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                         Altocrustacea
                                                           Miracrustacea
                                                             Superclass Hexapoda Latreille, 1825
                                                               Cercophora
                                                                 Class Insecta Linnaeus, 1758 - insects
                                                                   Subclass Dicondylia Haeckel, 1866
                                                                     Infraclass Pterygota Lang, 1888
                                                                       Metapterygota
                                                                         Neoptera Martynov, 1923
                                                                           Eumetabola
                                                                             Holometabola
                                                                               Order Coleoptera Linnaeus, 1758 - beetles
                                                                                 Suborder Polyphaga Emery, 1886
                                                                                   Series Cucujiformia Lameere, 1938
                                                                                     Superfamily Tenebrionoidea Latreille, 1802
                                                                                       Family Mordellidae Latreille, 1802
                                                                                         Genus Conalia Mulsant & Rey, 1858
                                                                                         Genus Glipa LeConte, 1859
                                                                                         Genus Glipodes LeConte, 1862
                                                                                         Genus Glipostenoda Ermisch, 1950
                                                                                         Genus Hoshihananomia Kono, 1935
                                                                                         Genus Isotrilophus Liljeblad, 1945
                                                                                         Genus Mordellistena Costa, 1854
                                                                                         Genus Paramordellaria Ermisch, 1968
                                                                                         Genus Tolidomordella Ermisch, 1950
                                                                                         Genus Tomoxia Costa, 1854
                                                                                         Genus Yakuhananomia Kono, 1935
                                                                                         1 Subfamily Ctenidiinae Franciscolo, 1951
                                                                                         2 Subfamily Mordellinaeᵀ Latreille, 1802