Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Dynastinae MacLeay, 1819
Genus Ancognatha Erichson, 1847
Genus Aphonides Rivers, 1889
Genus Aphonus LeConte, 1856
Genus Archophileurus Kolbe, 1910
Genus Aspidolea Bates, 1888
Genus Coscinocephalus Prell, 1936
Genus Dyscinetus Harold, 1869
Genus Euetheola Bates, 1888
Genus Golofa Hope, 1838
Genus Hemiphileurus Kolbe, 1910
Genus Ligyrus Burmeister, 1847
Genus Orizabus Fairmaire, 1878
Genus Podischnus Burmeister, 1847
Genus Stenocrates Burmeister, 1847
Genus Xyloryctes Hope, 1837
1 Tribe Hexodontini Lacordaire, 1856
2 Tribe Cyclocephalini Laporte, 1840
3 Tribe Dynastiniᵀ MacLeay, 1819
4 Tribe Oryctini Mulsant, 1842
5 Tribe Oryctoderini Endrödi, 1966
6 Tribe Pentodontini Mulsant, 1842
7 Tribe Agaocephalini Burmeister, 1847
8 Tribe Phileurini Burmeister, 1847