Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Obtectomera Minet, 1986, sedis mutabilis
Superfamily Papilionoidea Latreille, 1802, sedis mutabilis
Family Lycaenidae Leach, 1815 - gossamer-winged butterflies
Subfamily Poritiinae Doherty, 1886
Tribe Liptenini Röber, 1892
Subtribe Epitolina
01 Genus Aethiopana Bethune-Baker, 1915 [monotypic]
02 Genus Argyrotola Libert, 1999
03 Genus Batelusia Druce, 1910 [monotypic]
04 Genus Cephetola Libert, 1999
05 Genus Cerautola Libert, 1999
06 Genus Cercenia Libert, 1999
07 Genus Deloneura Trimen, 1868
08 Genus Epitolaᵀ Westwood, 1851
09 Genus Geritola Libert, 1999
10 Genus Hewitola Libert, 1999
11 Genus Hewitsonia Kirby, 1871
12 Genus Hypophytala Clench, 1965
13 Genus Iridana Aurivillius, 1920
14 Genus Neaveia Druce, 1910 [monotypic]
15 Genus Neoepitola Jackson, 1964 [monotypic]
16 Genus Phytala Westwood, 1851
17 Genus Powellana Bethune-Baker, 1908
18 Genus Pseudoneaveia Stempffer, 1964 [monotypic]
19 Genus Stempfferia Jackson, 1962
20 Genus Teratoneura Dudgeon, 1909 [monotypic]
21 Genus Tumerepedes Bethune-Baker, 1913 [monotypic]