Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Obtectomera Minet, 1986, sedis mutabilis
Superfamily Papilionoidea Latreille, 1802, sedis mutabilis
Family Lycaenidae Leach, 1815 - gossamer-winged butterflies
Subfamily Theclinae Swainson, 1830 (1820)
Tribe Eumaeini Doubleday, 1847
.1 Genus Beatheclus Bálint & Dahners, 2006
.2 Genus Cupathecla Bálint, 2005
.3 Genus Fasslantonius Bálint & Salazar, 2003
.4 Genus Suneve Bálint, 2006
0 [Genus Atara Zhdanko, 1996]
see Genus Rapala Moore, 1881
01 Section Eumaeus sectionᵀ
01.1.0 [Genus Epula Gistl, 1848]
see Genus Eumaeus Hübner, 1819
01.1.0 [Genus Eumaea Geyer, [1834]]
see Genus Eumaeus Hübner, 1819
01.1.0 [Genus Eumenia Godart, 1824]
see Genus Eumaeus Hübner, 1819
01.3.s [Genus Radissima Johnson, 1992]
see Genus Paiwarria Kaye, 1904
01.4.s [Genus Paraspiculatus Johnson & Constantino, 1997]
see Genus Mithras Hübner, 1818
02 Section Brangas section
02.3.0 [Genus Falerinota Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Enos Johnson, Kruse & Kroenlein, 1997
02.4.0 [Genus Cryptaenota Johnson, 1992]
see Genus Evenus Hübner, [1819]
02.4.0 [Genus Endymion Swainson, 1831]
see Genus Evenus Hübner, [1819]
02.4.0 [Genus Ipocia Brévignon, 2000]
see Genus Evenus Hübner, [1819]
02.4.0 [Genus Macusia Kaye, 1904]
see Genus Evenus Hübner, [1819]
02.4.0 [Genus Poetukulunma Brévignon, 2002]
see Genus Evenus Hübner, [1819]
02.5.0 [Genus "Annamaria" D'Abrera & Bálint, 2001]
see Genus Lamasina Robbins, 2002
03 Section Lamprospilus section
03.1.0 [Genus Gigantorubra Johnson, 1993]
see Genus Lamprospilus Geyer, 1832
03.2.0 [Genus Angulopis Johnson, 1991]
see Genus Electrostrymon Clench, 1961
03.2.0 [Genus Rubroserrata Johnson & Kroenlein, 1993]
see Genus Electrostrymon Clench, 1961
03.3.0 [Genus Antrissima Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Argentostriatus Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Calystryma Field, 1967]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Cyanodivida Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Distissima Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Femniterga Johnson, 1988]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Furcovalva Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Gigantofalca Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Klaufera Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Kroenleina Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Mercedes Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Morphissima Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Profieldia Johnson, 1992]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Reversustus Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Serratofalca Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Serratoterga Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Tergissima Johnson, 1988]
see Genus Calycopis Scudder, 1876
03.3.0 [Genus Terminospinissima Johnson, 1991]
see Genus Calycopis Scudder, 1876
03.6.0 [Genus Kisutam Johnson & Kroenlein, 1993]
see Genus Ziegleria Johnson, 1993
03.6.0 [Genus Pendantus Johnson & Kroenlein, 1993]
see Genus Ziegleria Johnson, 1993
04 Section Thereus section
04.1.0 [Genus Molus Hübner, [1819] [monotypic]]
see Genus Thereus Hübner, 1819
04.1.0 [Genus Noreena Johnson, MacPherson & Ingraham, 1986]
see Genus Thereus Hübner, 1819
04.1.0 [Genus Pedusa D'Abrera, 2001]
see Genus Thereus Hübner, 1819
04.1.0 [Genus Solanorum Johnson, 1992]
see Genus Thereus Hübner, 1819
04.1.0 [Genus Timokla Johnson, Kruse & Kroenlein, 1997]
see Genus Thereus Hübner, 1819
04.2.0 [Genus Heterosmaitia Clench, 1964]
see Genus Rekoa Kaye, 1904
04.3.0 [Genus Dolymorpha Holland, 1931 [monotypic]]
see Genus Arawacus Kaye, 1904
04.3.0 [Genus Polyniphes Kaye, 1904]
see Genus Arawacus Kaye, 1904
04.3.0 [Genus Tigrinota Johnson, 1992]
see Genus Arawacus Kaye, 1904
04.4.0 [Genus Orcya Johnson, 1990]
see Genus Contrafacia Johnson, 1989
05 Section Atlides section
05.1.0 [Genus Riojana D'Abrera & Bálint, 2001]
see Genus Atlides Hübner, [1819]
05.2.2.0 [Genus Aveexcrenota Salazar & Johnson, 1997]
see Genus Theritas Hübner, 1818
05.2.2.0 [Genus Denivia Johnson, 1992]
see Genus Theritas Hübner, 1818
05.2.2.0 [Genus Lucilda D'Abrera & Bálint, 2001]
see Genus Theritas Hübner, 1818
05.2.2.0 [Genus Margaritheclus Bálint, 2002]
see Genus Theritas Hübner, 1818
06 Section Allosmaitia section
06.2.0s1 [Genus Gibbossa Salazar & López, 2001]
see Genus Laothus Johnson, Kruse & Kroenlein, 1997
06.2.0s1o [Genus Gibbonota Salazar & López, 1996, nom. illeg.]
see Genus Gibbossa Salazar & López, 2001
07 Section Thestius section
08 Section Tmolus section
08.3.0 [Genus Crimsinota Johnson, 1993]
see Genus Nicolaea Johnson, 1993
08.3.0 [Genus Rindgea Johnson, 1993]
see Genus Nicolaea Johnson, 1993
08.7.0 [Genus Asymbiopsis Johnson & Le Crom, 1997]
see Genus Theclopsis Godman & Salvin, 1887
08.8.0 [Genus Decussata Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Ostrinotes Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Diminutina Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Dindyminotes Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Letizia Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Robustana Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Serratonotes Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Syedranota Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Treboniana Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
08.9.0 [Genus Zigirina Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Strephonota Johnson, Austin, Le Crom & Salazar, 1997
09 Section Panthiades section
09.1.0 [Genus Cycnus Hübner, 1819 [monotypic]]
see Genus Panthiades Hübner, [1819]
09.5.0 [Genus Eupsyche Scudder, 1876]
see Genus Parrhasius Hübner, 1819
10 Genus Strymon Hübner, 1818 - scrub hairstreaks
10.0 [Genus Callicista Grote, 1873]
see Genus Strymon Hübner, 1818
10.0 [Genus Eiseliana Toledo, 1978]
see Genus Strymon Hübner, 1818
10.0 [Genus Heoda Johnson, Miller & Herrera, 1992]
see Genus Strymon Hübner, 1818
11 Section Hypostrymon section
11.02.0 [Genus Sipaea Johnson, 1991]
see Genus Nesiostrymon Clench, 1964
11.02.0 [Genus Terra Johnson & Matusik, 1988]
see Genus Nesiostrymon Clench, 1964
11.08.0 [Genus Cyclotrichia Johnson, Austin, Le Crom & Salazar, 1997]
see Genus Celmia Johnson, 1991
11.08.0 [Genus Uzzia Johnson, 1991]
see Genus Celmia Johnson, 1991
11.09.0 [Genus Caerofethra Johnson, 1991]
see Genus Dicya Johnson, 1991
12 Section Satyrium section
12.1.0s01 [Genus Armenia Dubatolov & Korshunov, 1984]
see Genus Satyrium Scudder, 1876
12.1.0s02 [Genus Callipsyche Scudder, 1876 [monotypic]]
see Genus Satyrium Scudder, 1876
12.1.0s03 [Genus Chattendenia Tutt, 1908]
see Genus Satyrium Scudder, 1876
12.1.0s04 [Genus Euristrymon Clench, 1961]
see Genus Satyrium Scudder, 1876
12.1.0s05 [Genus Fixsenia Tutt, 1907]
see Genus Satyrium Scudder, 1876
12.1.0s06 [Genus Harkenclenus dos Passos, 1970 [monotypic]]
see Genus Satyrium Scudder, 1876
12.1.0s07 [Genus Necovatia Verity, 1951]
see Genus Satyrium Scudder, 1876
12.1.0s08 [Genus Neolycaena de Nicéville, 1890]
see Genus Satyrium Scudder, 1876
12.1.0s09 [Genus Nordmannia Tutt, 1907]
see Genus Satyrium Scudder, 1876
12.1.0s09o [Genus "Normannia"]
see Genus Nordmannia Tutt, 1907
12.1.0s10 [Genus Pseudothecla Strand, 1910 [monotypic]]
see Genus Satyrium Scudder, 1876
12.1.0s11 [Genus Rhymnaria Zhdanko, 1983]
see Genus Satyrium Scudder, 1876
12.1.0s12 [Genus Strymonidia Tutt, 1908]
see Genus Satyrium Scudder, 1876
12.1.0s12o [Genus "Strymondia" nom. null.]
see Genus Strymonidia Tutt, 1908
12.1.0s13 [Genus Superflua Strand, 1910 [monotypic]]
see Genus Satyrium Scudder, 1876
12.1.0s14 [Genus Thecliolia Strand, 1910 [monotypic]]
see Genus Satyrium Scudder, 1876
12.1.0s15o [Genus "Tuttiolia" Strand, 1910]
see Genus Tuttiola Strand, 1910
12.3.0 [Genus Arases Johnson, 1992]
see Genus Ocaria Clench, 1970
12.3.0 [Genus Galba Johnson, 1992, nom. illeg.]
see Genus Ocaria Clench, 1970
12.3.0 [Genus Lamasa Johnson, 1992]
see Genus Ocaria Clench, 1970
12.3.0 [Genus Variegatta Johnson, 1992]
see Genus Ocaria Clench, 1970
13 Section Erora section
13.1.0 [Genus Androcona Johnson, Eisele & MacPherson, 1993]
see Genus Erora Scudder, 1872
13.1.0 [Genus Necmitoura Johnson, Eisele & MacPherson, 1993]
see Genus Erora Scudder, 1872
13.1.0 [Genus Sarracenota Johnson, Eisele & MacPherson, 1993]
see Genus Erora Scudder, 1872
14 Section Callophrys section
14.1.0s01 [Genus Ahlbergia Bryk, 1946]
see Genus Callophrys Billberg, 1820
14.1.0s01o [Genus Ginzia Okano, 1947]
see Genus Ahlbergia Bryk, 1946
14.1.0s02 [Genus Cisincisalia Johnson, 1992]
see Genus Callophrys Billberg, 1820
14.1.0s03 [Genus Cissatsuma Johnson, 1992]
see Genus Callophrys Billberg, 1820
14.1.0s04 [Genus Deciduphagus Johnson, 1992]
see Genus Callophrys Billberg, 1820
14.1.0s05 [Genus Incisalia Scudder, 1872]
see Genus Callophrys Billberg, 1820
14.1.0s06 [Genus Loranthomitoura Ballmer & Pratt, 1992]
see Genus Callophrys Billberg, 1820
14.1.0s07 [Genus Mitoura Scudder, 1872]
see Genus Callophrys Billberg, 1820
14.1.0s08 [Genus Novosatsuma Johnson, 1992]
see Genus Callophrys Billberg, 1820
14.1.0s09 [Genus Sandia Clench & Ehrlich, 1960 [monotypic]]
see Genus Callophrys Billberg, 1820
14.1.0s10 [Genus Xamia Clench, 1961]
see Genus Callophrys Billberg, 1820
14.2.0 [Genus Antephrys Johnson, Eisele & MacPherson, 1993]
see Genus Cyanophrys Clench, 1961
14.2.0 [Genus Apophrys Johnson & Le Crom, 1997]
see Genus Cyanophrys Clench, 1961
14.2.0 [Genus Mesocyanophrys Johnson & Le Crom, 1997]
see Genus Cyanophrys Clench, 1961
14.2.0 [Genus Plesiocyanophrys Johnson, Eisele & MacPherson, 1993]
see Genus Cyanophrys Clench, 1961
15 Section Micandra section
15.01.0 [Genus Egides Johnson, Kruse & Kroenlein, 1997]
see Genus Micandra Staudinger, 1888
15.03.0 [Genus Owda Johnson, Kruse & Kroenlein, 1997]
see Genus Johnsonita Salazar & Constantino, 1995
15.04.0 [Genus Bussa Johnson, Kruse & Kroenlein, 1997, nom. illeg.]
see Genus Brevianta Johnson, Kruse & Kroenlein, 1997
15.05.0 [Genus Jagiello Bálint & Wojtusiak, 2000]
see Genus Timaeta Johnson, Kruse & Kroenlein, 1997
15.05.0 [Genus Trochusinus Johnson, Salazar & Vélez, 1997]
see Genus Timaeta Johnson, Kruse & Kroenlein, 1997
15.06.0 [Genus Paralustrus Johnson, 1992]
see Genus Rhamma Johnson, 1992
15.06.0 [Genus Pontirama Johnson, 1992]
see Genus Rhamma Johnson, 1992
15.06.0 [Genus Shapiroana Johnson, 1992]
see Genus Rhamma Johnson, 1992
15.10.0 [Genus Abloxurina Johnson, 1992]
see Genus Penaincisalia Johnson, 1990
15.10.0 [Genus Candora Johnson, 1992]
see Genus Penaincisalia Johnson, 1990
15.10.0 [Genus Pons Johnson, 1992]
see Genus Penaincisalia Johnson, 1990
15.10.0 [Genus Thecloxurina Johnson, 1992]
see Genus Penaincisalia Johnson, 1990