Natura - nature
  
 actualia - actual entities
    
 Mundus Plinius - physical world
      
 naturalia - natural bodies
        
 Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
          
 Clade neomura Cavalier-Smith, 1987
            
 Domain Eukaryota Chatton, 1925 - eukaryotes
              
 Clade discaria Cavalier-Smith, 2021
                
 Clade dorsates Cavalier-Smith, 2021
                  
 Clade podiates Cavalier-Smith, 2012
                    
 Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                      
 Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                        
 Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                          
 Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                            
 Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
                              
 Kingdom Animalia Linnaeus, 1758 - animals
                                
 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                  
 Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                    
 Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                      
 Infrakingdom Protostomia Grobben, 1908
                                        
 Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                          
 Panarthropoda Nielsen, 1995 - panarthropods
                                            
 Tactopoda Budd, 2001
                                              
 Phylum Arthropoda Siebold, 1848 - arthropods
                                                
 Euarthropoda
                                                  
 Antennulata
                                                    
 Subphylum Mandibulata Snodgrass, 1938
                                                      
 Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                        
 Altocrustacea
                                                          
 Miracrustacea
                                                            
 Superclass Hexapoda Latreille, 1825
                                                              
 Cercophora
                                                                
 Class Insecta Linnaeus, 1758 - insects
                                                                  
 Subclass Dicondylia Haeckel, 1866
                                                                    
 Infraclass Pterygota Lang, 1888
                                                                      
 Metapterygota
                                                                        
 Neoptera Martynov, 1923
                                                                          
 Eumetabola
                                                                            
 Holometabola
                                                                              
 Order Coleoptera Linnaeus, 1758 - beetles
                                                                                
 Suborder Polyphaga Emery, 1886
                                                                                  
 Series Cucujiformia Lameere, 1938
                                                                                    
 Superfamily Tenebrionoidea Latreille, 1802
                                                                                      
 01 Family Mycetophagidae Leach, 1815
                                                                                      
 02 Family Archeocrypticidae Kaszab, 1964
                                                                                      
 03 Family Pterogeniidae Crowson, 1953
                                                                                      
 04 Family Ciidae Leach, 1819
                                                                                      
 05 Family Tetratomidae Billberg, 1820
                                                                                      
 06 Family Melandryidae Leach, 1815
                                                                                      
 07 Family Mordellidae Latreille, 1802
                                                                                      
 08 Family Ripiphoridae Gemminger, 1870 (1853)
                                                                                      
 09 Family Zopheridae Solier, 1834
                                                                                      
 10 Family Ulodidae Pascoe, 1869
                                                                                      
 11 Family Promecheilidae Lacordaire, 1859
                                                                                      
 12 Family Chalcodryidae Watt, 1974
                                                                                      
 13 Family Trachelostenidae Lacordaire, 1859
                                                                                      
 14 Family Tenebrionidaeᵀ Latreille, 1802 - darkling beetles
                                                                                      
 15 Family Prostomidae Thomson, 1859
                                                                                      
 16 Family Synchroidae Lacordaire, 1859
                                                                                      
 17 Family Stenotrachelidae Thomson, 1859
                                                                                      
 18 Family Oedemeridae Latreille, 1810
                                                                                      
 19 Family Meloidae Gyllenhal, 1810 - blister beetles
                                                                                      
 20 Family Mycteridae Oken, 1843
                                                                                      
 21 Family Boridae Thomson, 1859
                                                                                      
 22 Family Trictenotomidae Blanchard, 1845
                                                                                      
 23 Family Pythidae Solier, 1834
                                                                                      
 24 Family Pyrochroidae Latreille, 1807
                                                                                      
 25 Family Salpingidae Leach, 1815
                                                                                      
 26 Family Anthicidae Latreille, 1819
                                                                                      
 27 Family Aderidae Winkler, 1927
                                                                                      
 28 Family Scraptiidae Gistel, 1848