Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 Clade neozoa Cavalier-Smith 1993
                   Clade scotokaryotes Cavalier-Smith 2000
                     Clade podiates Cavalier-Smith 2012
                       [crown] Clade Amorphea Adl et al. 2012
                         [crown] Clade Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                       Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                             Tactopoda Budd, 2001
                                               Phylum Arthropoda Siebold, 1848 - arthropods
                                                 Euarthropoda
                                                   Antennulata
                                                     Subphylum Mandibulata Snodgrass, 1938
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                         Altocrustacea
                                                           Miracrustacea
                                                             Superclass Hexapoda Latreille, 1825
                                                               Cercophora
                                                                 Class Insecta Linnaeus, 1758 - insects
                                                                   Subclass Dicondylia Haeckel, 1866
                                                                     Infraclass Pterygota Lang, 1888
                                                                       Metapterygota
                                                                         Neoptera Martynov, 1923
                                                                           Eumetabola
                                                                             Holometabola
                                                                               Superorder Panorpida
                                                                                 Amphiesmenoptera
                                                                                   Order Lepidoptera Linnaeus, 1758 - butterflies, moths
                                                                                     Suborder Glossata Fabricius, 1775
                                                                                       Coelolepida Nielsen & Kristensen, 1996
                                                                                         Myoglossata Kristensen & Nielsen, 1981
                                                                                           Neolepidoptera Packard, 1895
                                                                                             Infraorder Heteroneura Tillyard, 1918
                                                                                               Eulepidoptera Kiriakoff, 1948
                                                                                                 Ditrysia Börner, 1925
                                                                                                   Apoditrysia Minet, 1983, sedis mutabilis
                                                                                                     Obtectomera Minet, 1986, sedis mutabilis
                                                                                                       Superfamily Papilionoidea Latreille, 1802, sedis mutabilis
                                                                                                         Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                                                                                                           Subfamily Cyrestinae Guenée, 1865
                                                                                                             1 Tribe Cyrestiniᵀ Guenée, 1865
                                                                                                             2 Tribe Pseudergolini Jordan, 1898