Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Scarabaeinaeᵀ Latreille, 1802
Tribe Onthophagini Burmeister, 1846
Genus Onthophagusᵀ Latreille, 1802
Onthophagus acuminatus Harold, 1880
Onthophagus aeruginosus Roth, 1851
Onthophagus alcyonides D'Orbigny, 1913
Onthophagus asperulus D'orbigny, 1905
Onthophagus australis Guerin, 1830
Onthophagus beesoni Arrow, 1931
Onthophagus binodis Thunberg, 1818
Onthophagus capella Kirby, 1818
Onthophagus clypeatus Blanchard, 1843
Onthophagus cochisus Brown, 1927
Onthophagus coscineus Bates, 1887
Onthophagus cribripennis D'orbigny, 1902
Onthophagus crinitus Harold, 1869
Onthophagus ensifer Boucomont, 1914
Onthophagus evanidus Harold, 1869
Onthophagus ferox Harold, 1867
Onthophagus fuliginosus Erichson, 1842
Onthophagus gazella Heyden, 1913
Onthophagus granulatus Boheman, 1858
Onthophagus haagi Harold, 1867
Onthophagus haematopus Harold, 1875
Onthophagus hecate (Panzer, 1794)
Onthophagus incensus Say, 1837
Onthophagus knulli Howden & Cartwright, 1963
Onthophagus laminatus Macleay, 1864
Onthophagus lanista Gerstaecker, 1873
Onthophagus marginicollis Harold, 1880
Onthophagus mjobergi Gillet, 1925
Onthophagus muticus Macleay, 1864
Onthophagus nigriventris D'orbigny, 1905
Onthophagus nuchicornis (C. Linnaeus, 1758)
Onthophagus orpheus (Panzer, 1794)
Onthophagus pennsylvanicus Harold, 1871
Onthophagus pentacanthus Harold, 1867
Onthophagus praecellens Bates, 1887
Onthophagus raffrayi Harold, 1886
Onthophagus rana Arrow, 1931
Onthophagus rangifer Klug, 1855
Onthophagus sagittarius (Fabricius, 1775)
Onthophagus sharpi Harold, 1875
Onthophagus sloanei Blackburn, 1903
Onthophagus stockwelli Howden & Young, 1981
Onthophagus sugillatus Klug, 1855
Onthophagus taurus Schreber, 1759
Onthophagus tersidorsis D'Orbigny, 1902
Onthophagus vermiculatus Frey, 1970
Onthophagus xanthomerus Bates, 1887