Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 neozoa Cavalier-Smith 1993
                   scotokaryotes Cavalier-Smith 2000
                     podiates Cavalier-Smith 2012 [nom. cl. nov.]
                       Amorphea Adl et al. 2012 [nom. cl. nov.]
                         Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
                             Kingdom Fungi R.T. Moore (1980) - fungi
                               Subkingdom Dikarya Hibbett, T.Y. James & Vilgalys (2007)
                                 Phylum Basidiomycota R.T. Moore (1980) - basidiomycetes
                                   Subphylum Agaricomycotina Doweld (2001)
                                     Class Agaricomycetesᵀ Doweld (2001)
                                       Subclass Agaricomycetidaeᵀ Locq. ex Parmasto (1986)
                                         Order Boletales E.-J. Gilbert (1931)
                                           1 Genus Corditubera Henn. (1897), incertae sedis
                                           2 Genus Corneromyces Ginns (1976), incertae sedis
                                           3 Genus Marthanella States & Fogel (1999), incertae sedis
                                           4 Genus Phaeoradulum Pat. (1900), incertae sedis
                                           5? Family Protogastraceae Zeller (1934) [monotypic]
                                           6? Family Rhizopogonaceae Gäum. & C.W. Dodge (1928)
                                           7 Family Tapinellaceae C. Hahn (1999)
                                           8.1.1 Family Hygrophoropsidaceae Kühner (1980)
                                           8.1.2 Family Serpulaceae Jarosch & Bresinsky (2001)
                                           8.2.1 Family Coniophoraceae Ulbr. (1928)
                                           8.2.2.1.1 Family Gomphidiaceae Maire ex Jülich (1982)
                                           8.2.2.1.2 Family Suillaceae Besl & Bresinsky (1997)
                                           8.2.2.2.1.1 Family Paxillaceae Lotsy (1907)
                                           8.2.2.2.1.2.1 Family Boletinellaceae P.M. Kirk, P.F. Cannon & J.C. David (2001)
                                           8.2.2.2.1.2.2.1 Family Gyroporaceae Manfr. Binder & Bresinsky (2002) [monotypic]
                                           8.2.2.2.1.2.2.2.1 Family Calostomataceae E. Fisch. (1900) [monotypic]
                                           8.2.2.2.1.2.2.2.2 Family Sclerodermataceae Corda (1842)
                                           8.2.2.2.1.2.2.2.3? Family "Diplocystidiaceae" Kreisel (1974), orth. inval.
                                           8.2.2.2.2 Family Boletaceaeᵀ Chevall. (1826)
                                           9? Family Gasterellaceae Zeller (1948) [monotypic]