Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Obtectomera Minet, 1986, sedis mutabilis
Superfamily Papilionoidea Latreille, 1802, sedis mutabilis
Family Lycaenidae Leach, 1815 - gossamer-winged butterflies
Subfamily Theclinae Swainson, 1830 (1820)
Tribe Thecliniᵀ Swainson, 1830 (1820)
Genus Amblopala Leech, 1893 [monotypic]
Genus Antigius Sibatani & Ito, 1942
Genus Araragi Sibatani & Ito, 1942 [monotypic]
Genus Artopoetes Chapman, 1909 [monotypic]
Genus Austrozephyrus Howarth, 1957 [monotypic]
Genus Borneozephyrus Koiwaya, 2007
Genus Chaetoprocta Nicéville, 1890 [monotypic]
Genus Cordelia Shirôzu & Yamamoto, 1956 [monotypic]
Genus Coreana Tutt, 1907
Genus Esakiozephyrus Shirôzu & Yamamoto, 1956
Genus Fujiokaozephyrus Koiwaya, 2007
Genus Goldia Dubatolov & Korshunov, 1990
Genus Gonerilia Shirôzu & Yamamoto, 1956 [monotypic]
Genus Habrodais Scudder, 1876 [monotypic]
Genus Hayashikeia Fujioka, 2003
Genus Howarthia Shirôzu & Yamamoto, 1956 [monotypic]
Genus Inomataozephyrus Koiwaya, 2007
Genus Iozephyrus Wang & Fan, 2002
Genus Iratsume Sibatani & Ito, 1942 [monotypic]
Genus Iwaseozephyrus Fujioka, 1994
Genus Kameiozephyrus Koiwaya, 2007
Genus Kawazoeozephyrus Koiwaya, 2007
Genus Leucantigius Shirôzu & Murayama, 1951 [monotypic]
Genus Nanlingozephyrus Wang & Pang, 1998
Genus Neogonerilia Koiwaya, 2007
Genus Noseozephyrus Koiwaya, 2007
Genus Palawanozephyrus Koiwaya, 2007
Genus Protantigius Shirôzu & Yamamoto, 1956 [monotypic]
Genus Proteuaspa Koiwaya & Morita, 2003
Genus Pseudogonerilia Koiwaya, 2007
Genus Ravenna Shirôzu & Yamamoto, 1956 [monotypic]
Genus Saigusaozephyrus Koiwaya, 1993
Genus Shaanxiana Koiwaya, 1993
Genus Shirozua Sibatani & Ito, 1942 [monotypic]
Genus Shirozuozephyrus Koiwaya, 2007
Genus Shizuyaozephyrus Koiwaya, 2003
Genus Sibataniozephyrus Inomata, 1986
Genus Tasogare Saigusa, 1993
Genus Teratozephyrus Sibitani, 1946
Genus Thermozephyrus Inomata & Itagaki, 1986
Genus Uedaozephyrus Koiwaya, 2007
Genus Wagimo Sibatani & Ito, 1942
Genus Yamamotozephyrus Saigusa, 1993
Genus Yuhbae Saigusa, 1993
1 Genus Laeosopis Rambur, 1858
2 Genus Theclaᵀ Fabricius, 1807
3 Genus Ussuriana Tutt, 1907
4 Genus Japonica Tutt, 1907 [monotypic]
5 Genus Hypaurotis Scudder, 1876
6 Genus Euaspa Moore, 1884 [monotypic]
7 Genus Chrysozephyrus Shirôzu & Yamamoto, 1956
8 Genus Favonius Sibatani & Ito, 1942
9.1 Genus Neozephyrus Sibatani & Ito, 1942
9.2 Genus Quercusia Verity, 1943 [monotypic]