Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 neozoa Cavalier-Smith 1993
                   scotokaryotes Cavalier-Smith 2000
                     podiates Cavalier-Smith 2012 [nom. cl. nov.]
                       Amorphea Adl et al. 2012 [nom. cl. nov.]
                         Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
                                 Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004 [nom. cl. nov.]
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002 [nom. cl. nov.]
                                       Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                             Phylum Arthropoda von Siebold, 1848 - arthropods
                                               Euarthropoda
                                                 Antennulata
                                                   Subphylum Mandibulata Snodgrass, 1938
                                                     Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                       Altocrustacea
                                                         Miracrustacea
                                                           Superclass Hexapoda Latreille, 1825
                                                             Class Insecta Linnaeus, 1758 - insects
                                                               Subclass Dicondylia
                                                                 Infraclass Pterygota
                                                                   Metapterygota
                                                                     Neoptera
                                                                       Eumetabola
                                                                         Holometabola
                                                                           Superorder Panorpida
                                                                             Amphiesmenoptera
                                                                               Order Lepidoptera Linnaeus, 1758 - butterflies, moths
                                                                                 Suborder Glossata Fabricius, 1775
                                                                                   Coelolepida Nielsen & Kristensen, 1996
                                                                                     Myoglossata Kristensen & Nielsen, 1981
                                                                                       Neolepidoptera Packard, 1895
                                                                                         Infraorder Heteroneura Tillyard, 1918
                                                                                           Eulepidoptera Kiriakoff, 1948
                                                                                             Ditrysia Börner, 1925
                                                                                               Apoditrysia Minet, 1983, sedis mutabilis
                                                                                                 Obtectomera Minet, 1986, sedis mutabilis
                                                                                                   Superfamily Papilionoidea Latreille, 1802, sedis mutabilis
                                                                                                     Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                                                                                                       Subfamily Satyrinae Boisduval, 1833
                                                                                                         Tribe Zetherini Reuter, 1896
                                                                                                           1.1 Genus Callarge Leech, 1892
                                                                                                           1.2 Genus Penthema Doubleday, 1848
                                                                                                           1.3 Genus Zetheraᵀ Felder, 1861
                                                                                                           2.1 Genus Ethope Moore, 1866
                                                                                                           2.2 Genus Neorina Westwood, 1850