Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Genus Bolbogonium Boucomont, 1911
Genus Bolbohamatum Krikken, 1980
Genus Charitovalgus Kolbe, 1904
Genus Clinterocera Motschoulsky, 1857[1858]
Genus Heterovalgus Krikken, 1978
Genus Ingrisma Fairmaire, 1893
Genus Ischiopsopha Gestro, 1874
Genus Mecinonota Kraatz, 1892
Genus Microlomaptera Kraatz, 1885
Genus Mimobolbus Vulcano, Martinez & Pereira, 1969
Genus Oncosterna Thomson, 1880
Genus Peotoxus Krikken, 1983
Genus Platygeniops Krikken, 1978
Genus Plectrone Wallace, 1867
Genus Rhabdotops Krikken, 1981
Genus Rhinacosmus Kraatz, 1895
Genus Ruteraetia Krikken, 1980
Genus Synapsis Bates, 1868
Genus Termitodius Wasmann, 1894
01 Family Pleocomidae LeConte, 1861
02 Family Geotrupidae Latreille, 1802
03 Family Belohinidae Paulian, 1959
04 Family Passalidae Leach, 1815
05.1 Family Trogidae MacLeay, 1819
05.2 Family Glaresidae Kolbe, 1905
06 Family Diphyllostomatidae Holloway, 1972
07 Family Lucanidae Latreille, 1804 - stag beetles
08 Family Ochodaeidae Mulsant & Rey, 1871
09.1 Family Hybosoridae Erichson, 1847
09.2 Family Glaphyridae MacLeay, 1819
09.3 Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
10 Ichnofamily †Coprinisphaeridae Genise, 2004
11 Ichnofamily †Pallichnidae Genise, 2004